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Kouphichnium
''Kouphichnium'' is an ichnogenus that has been attributed to limulids (horseshoe crabs). ''Kouphichnium'' fossils resemble the footprints of birds, sometimes in conjunction with a medial line, and were initially thought to be bird or pterosaur tracks. The footprints are now believed to be the imprints of specialized limulid appendages, called pushers, terminating in four plates, used to push against the sediment. The medial line is left by the animal's telson. This ichnogenus is registered in the Carboniferous to the Cretaceous, in marine marginal environments in Tennessee, the United States and in Poland, as well as non-marine environments of Argentina and England, among others.Aceñolaza, F. G., & Buatois, L. A. (1991). Trazas fósiles del Paleozoico superior continental argentino. Ameghiniana', 28, 89-108. The genus contains five species, ''K. arizonae'', ''K. cordifomnis'', ''K. lithographicum'', ''K. minusculum'' and ''K. walchi''. See also * List of xiphosuran genera * ...
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Mesolimulus
''Mesolimulus'' is an extinct genus of horseshoe crab. The best known examples are found in Solnhofen limestone near Solnhofen, Bavaria, Germany. Originally assigned to the living genus ''Limulus'', they are related to and look virtually identical to modern horseshoe crabs. Other species assigned to ''Mesolimulus'' have been recorded spanning over 140 million years from the Middle Triassic to Late Cretaceous from England, Spain, Siberia and Morocco. Kouphichnium The unusual fossil trackway, trackways left by these animals at Solnhofen were originally thought to have been made by birds or pterosaurs, since they included a cross-shaped marking comparable with the impression of a typical bird foot. Eventually they were correctly identified as arthropod footprints and named ''Kouphichnium walchi'', as proved by some remarkable 'death traces' showing a ''Mesolimulus'' circling round on itself before dying. The last leg of more advanced horseshoe crabs is modified into a so-called 'p ...
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List Of Xiphosuran Genera
This list of xiphosurans is a comprehensive listing of all genera that have ever been included in the order Xiphosura, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (''nomen dubium''), or were not formally published (''nomen nudum''), as well as Synonym (zoology), junior synonyms of more established names, genera that are no longer considered xiphosurans and misspellings. The list currently includes 54 names out of which 31 are considered valid xiphosuran genera. As of 2019, there were around 80 extinct species of xiphosurans and 4 extant species of xiphosurans recognized as valid. Naming conventions and terminology There is no "official" or "canonical" list of xiphosuran genera. The closest thing is found contained in the regularly updated ''Summary list of fossil spiders and their relatives'' in the World Spider Catalog, which include just the extinct species. The vast majority or all ...
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Mortichnia
A mortichnia is the "death march", or last walk, of a living creature. These are sometimes preserved as fossil footprints. Notable examples In 2002 the mortichnia of a horseshoe crab was found in lithographic limestone in Bavaria, Germany. The trail measured 9.7m and was left about 150 million years ago when the crab died in an anoxic lagoon. The footprints left enough evidence for researchers to determine that the creature probably fell into the lagoon upside-down, righted itself, and started walking before succumbing to the anoxic conditions of the water. The trackway is currently exhibited at the Wyoming Dinosaur Center. See also * ''Kouphichnium'', an ichnogenus An ichnotaxon (plural ichnotaxa) is "a taxon based on the fossilized work of an organism", i.e. the non-human equivalent of an artifact. ''Ichnotaxa'' comes from the Greek ίχνος, ''ichnos'' meaning ''track'' and ταξις, ''taxis'' meaning ... attributed to horseshoe crabs References Walking Anoxic wate ...
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Limulid
Horseshoe crabs are marine and brackish water arthropods of the family Limulidae and the only living members of the order Xiphosura. Despite their name, they are not true crabs or crustaceans: they are chelicerates, most closely related to arachnids, such as spiders and scorpions. Horseshoe crabs live primarily in and around shallow coastal waters on soft, sandy or muddy bottoms. They are generally found in the intertidal zone at spring high tides. They are eaten in some parts of Asia, and used as fishing bait, in fertilizer and in science (especially ''Limulus'' amebocyte lysate). In recent years, population declines have occurred as a consequence of coastal habitat destruction and overharvesting. Tetrodotoxin may be present in one horseshoe crab species, ''Carcinoscorpius rotundicauda''. Fossil records for horseshoe crabs extend back as far as 480 million years ago, with extant forms being living fossils. A 2019 molecular analysis places them as the sister group of ...
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Footprint
Footprints are the impressions or images left behind by a person walking or running. Hoofprints and pawprints are those left by animals with hooves or paws rather than feet, while "shoeprints" is the specific term for prints made by shoes. They may either be indentations in the ground or something placed onto the surface that was stuck to the bottom of the foot. A "trackway" is a set of footprints in soft earth left by a life-form; animal tracks are the footprints, hoofprints, or pawprints of an animal. Footprints can be followed when tracking during a hunt or can provide evidence of activities. Some footprints remain unexplained, with several famous stories from mythology and legend. Others have provided evidence of prehistoric life and behaviours. Footprints in detective work The print left behind at a crime scene can give vital evidence to the perpetrator of the crime. Shoes have many different prints based on the sole design and the wear that it has received – this can ...
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Pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Appendage
An appendage (or outgrowth) is an external body part, or natural prolongation, that protrudes from an organism's body. In arthropods, an appendage refers to any of the homologous body parts that may extend from a body segment, including antennae, mouthparts (including mandibles, maxillae and maxillipeds), gills, locomotor legs ( pereiopods for walking, and pleopods for swimming), sexual organs (gonopods), and parts of the tail (uropods). Typically, each body segment carries one pair of appendages. An appendage which is modified to assist in feeding is known as a maxilliped or gnathopod. In vertebrates, an appendage can refer to a locomotor part such as a tail, fins on a fish, limbs (legs, flippers or wings) on a tetrapod; exposed sex organ; defensive parts such as horns and antlers; or sensory organs such as auricles, proboscis ( trunk and snout) and barbels. Appendages may become ''uniramous'', as in insects and centipedes, where each appendage comprises a single ...
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Telson
The telson () is the posterior-most division of the body of an arthropod. Depending on the definition, the telson is either considered to be the final segment of the arthropod body, or an additional division that is not a true segment on account of not arising in the embryo from teloblast areas as other segments. It never carries any appendages, but a forked "tail" called the caudal furca may be present. The shape and composition of the telson differs between arthropod groups. Crustaceans In lobsters, shrimp and other decapods, the telson, along with the uropods, forms the tail fan. This is used as a paddle in the caridoid escape reaction ("lobstering"), whereby an alarmed animal rapidly flexes its tail, causing it to dart backwards. Krill can reach speeds of over 60 cm per second by this means. The trigger time to optical stimulus is, in spite of the low temperatures, only 55 milliseconds. In the Isopoda and Tanaidacea (superorder Peracarida), the last abdominal b ...
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Carboniferous
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferous'' means "coal-bearing", from the Latin '' carbō'' ("coal") and '' ferō'' ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern 'system' names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession. The Carboniferous is often treated in North America as two geological periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial animal life was well established by the Carboniferous Period. Tetrapods (four limbed vertebrates), which had originated from lobe-finned fish during the preceding Devonian, became pentadactylous in and diversified during the Carboniferous, including early amphibian line ...
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Fossil Trackway
A fossil track or ichnite (Greek "''ιχνιον''" (''ichnion'') – a track, trace or footstep) is a fossilized footprint. This is a type of trace fossil. A fossil trackway is a sequence of fossil tracks left by a single organism. Over the years, many ichnites have been found, around the world, giving important clues about the behaviour (and foot structure and stride) of the animals that made them. For instance, multiple ichnites of a single species, close together, suggest 'herd' or 'pack' behaviour of that species. Combinations of footprints of different species provide clues about the interactions of those species. Even a set of footprints of a single animal gives important clues, as to whether it was bipedal or quadrupedal. In this way, it has been suggested that some pterosaurs, when on the ground, used their forelimbs in an unexpected quadrupedal action. Special conditions are required, in order to preserve a footprint made in soft ground (such as an alluvial plain or a f ...
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Tennessee
Tennessee ( , ), officially the State of Tennessee, is a landlocked state in the Southeastern region of the United States. Tennessee is the 36th-largest by area and the 15th-most populous of the 50 states. It is bordered by Kentucky to the north, Virginia to the northeast, North Carolina to the east, Georgia, Alabama, and Mississippi to the south, Arkansas to the southwest, and Missouri to the northwest. Tennessee is geographically, culturally, and legally divided into three Grand Divisions of East, Middle, and West Tennessee. Nashville is the state's capital and largest city, and anchors its largest metropolitan area. Other major cities include Memphis, Knoxville, Chattanooga, and Clarksville. Tennessee's population as of the 2020 United States census is approximately 6.9 million. Tennessee is rooted in the Watauga Association, a 1772 frontier pact generally regarded as the first constitutional government west of the Appalachian Mountains. Its name derives from "Tanas ...
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United States
The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territories, nine Minor Outlying Islands, and 326 Indian reservations. The United States is also in free association with three Pacific Island sovereign states: the Federated States of Micronesia, the Marshall Islands, and the Republic of Palau. It is the world's third-largest country by both land and total area. It shares land borders with Canada to its north and with Mexico to its south and has maritime borders with the Bahamas, Cuba, Russia, and other nations. With a population of over 333 million, it is the most populous country in the Americas and the third most populous in the world. The national capital of the United States is Washington, D.C. and its most populous city and principal financial center is New York City. Paleo-Americ ...
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