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Kerberosaurus
''Kerberosaurus'' (meaning " Kerberos lizard") was a genus of saurolophine duckbill dinosaur from the late Maastrichtian-age Upper Cretaceous Tsagayan Formation of Blagoveshchensk, Amur Region, Russia (dated to 66 million years ago).Godefroit, P., Lauters, P., Van Itterbeeck, J., Bolotsky, Y. and Bolotsky, I.Y. (2011). "Recent advances on study of hadrosaurid dinosaurs in Heilongjiang (Amur) River area between China and Russia." ''Global Geology'', 2011(3). It is based on bonebed material including skull remains indicating that it was related to ''Saurolophus'' and ''Prosaurolophus''. History In 1984, Yuri Bolotsky and the Amur Complex Integrated Research Institute discovered a large dinosaur bonebed at Blagoveschensk. Most of the remains were of ''Amurosaurus'' (a lambeosaurine hadrosaur), but some came from turtles, crocodilians, theropods, nodosaurids, and a new hadrosaurine. For the hadrosaurine, cranial material (holotype AENM 1/319, braincase, plus others) was dist ...
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Kundurosaurus
''Kundurosaurus'' is an extinct genus of saurolophine hadrosaurid dinosaur known from the Latest Cretaceous (probably Late Maastrichtian stage) of Amur Region, Far Eastern Russia. It contains a single species, ''Kundurosaurus nagornyi''. Description ''Kundurosaurus'' was a medium-sized hadrosaur, measuring long and weighing . It is a saurolophine diagnosed by four autapomorphies, unique derived traits. It has a prominent and thick ridge on the lateral side of the nasal that borders caudally the circumnasal depression and invades the caudal plate of the nasal. Its caudal buttress of the proximal head of the scapula is oriented quite laterally, parallel to the pseudoacromial process. The preacetabular process of the ilium is straight and only moderately deflected ventrally by an angle of 160°. It does not reach the level of the plane formed by the bases of the iliac and pubic peduncles. Finally, with ''Kundurosaurus'' the axis of the postacetabular process of the ilium is stron ...
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Hadrosaur
Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which includes genera such as ''Edmontosaurus'' and ''Parasaurolophus'', was a common group of herbivores during the Late Cretaceous Period. Hadrosaurids are descendants of the Upper Jurassic/Lower Cretaceous iguanodontian dinosaurs and had a similar body layout. Hadrosaurs were among the most dominant herbivores during the Late Cretaceous in Asia and North America, and during the close of the Cretaceous several lineages dispersed into Europe, Africa, South America and Antarctica. Like other ornithischians, hadrosaurids had a predentary bone and a pubic bone which was positioned backwards in the pelvis. Unlike more primitive iguanodonts, the teeth of hadrosaurids are stacked into complex structures known as dental batteries, which acted as effective g ...
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Saurolophinae
Saurolophinae is a subfamily of hadrosaurid dinosaurs. It has since the mid-20th century generally been called the Hadrosaurinae, a group of largely non-crested hadrosaurs related to the crested sub-family Lambeosaurinae. However, the name Hadrosaurinae is based on the genus ''Hadrosaurus'' which was found in more recent studies to be more primitive than either lambeosaurines or other traditional "hadrosaurines", like ''Edmontosaurus'' and '' Saurolophus''. As a result of this, the name Hadrosaurinae was dropped or restricted to ''Hadrosaurus'' alone, and the subfamily comprising the traditional "hadrosaurines" was renamed the Saurolophinae. Recent phylogenetic work by Hai Xing indicates that ''Hadrosaurus'' is placed within the monophyletic group containing all non-lambeosaurine hadrosaurids. Under this view, the traditional Hadrosaurinae is resurrected, with the Hadrosauridae being divided into two clades: Hadrosaurinae and Lambeosaurinae. Saurolophinae was first defined as ...
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Amurosaurus
''Amurosaurus'' (; "Amur lizard") is a genus of lambeosaurine hadrosaurid dinosaur found in the latest Cretaceous period (66 million years ago)Godefroit, P., Lauters, P., Van Itterbeeck, J., Bolotsky, Y. and Bolotsky, I.Y. (2011). "Recent advances on study of hadrosaurid dinosaurs in Heilongjiang (Amur) River area between China and Russia." ''Global Geology'', 2011(3). of eastern Asia. Fossil bones of adults are rare, but an adult would most likely have been at least long. According to Gregory S. Paul, it was about long and weighed about . Discovery and naming Russian paleontologists Yuri Bolotsky and Sergei Kurzanov first described and named this dinosaur in 1991. The generic name is derived from the Amur River and the Greek word ''sauros'' ("lizard"). The Amur (called Heilongjiang or "Black Dragon River" in Chinese) forms the border of Russia and China, and is near where this dinosaur's remains were found. There is one known species (''A. riabinini''), named in honor of ...
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Saurolophus
''Saurolophus'' (; meaning "lizard crest") is a genus of large hadrosaurid dinosaur from the Late Cretaceous period of Asia and North America, that lived in what is now the Horseshoe Canyon and Nemegt formations about 70 million to 68 million years ago. It is one of the few genera of dinosaurs known from multiple continents. The type species, ''S. osborni'', was described by Barnum Brown in 1912 from Canadian fossils. A second valid species, ''S. angustirostris'', is represented by numerous specimens from Mongolia, and was described by Anatoly Konstantinovich Rozhdestvensky. ''Saurolophus'' is distinguished by a spike-like crest which projects up and back from the skull. It was a herbivorous dinosaur which could move about either bipedally or quadrupedally. Discovery and history Barnum Brown recovered the first described remains of ''Saurolophus'' in 1911, including a nearly complete skeleton (AMNH 5220). Now on display in the American Museum of Natural History, this skeleto ...
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Prosaurolophus
''Prosaurolophus'' (; meaning "before ''Saurolophus''", in comparison to the later dinosaur with a similar head crest) is a genus of hadrosaurid (or duck-billed) dinosaur from the Late Cretaceous of North America. It is known from the remains of at least 25 individuals belonging to two species, including skulls and skeletons, but it remains obscure. Its fossils have been found in the late Campanian-age Upper Cretaceous Dinosaur Park Formation in Alberta, and the roughly contemporaneous Two Medicine Formation in Montana, dating to around 75.5-74.0 million years ago. Its most recognizable feature is a small solid crest formed by the nasal bones, sticking up in front of the eyes. The type species is ''P. maximus'', described by American paleontologist Barnum Brown of the American Museum of Natural History in 1916. A second species, ''P. blackfeetensis'', was described by Jack Horner of the Museum of the Rockies in 1992. The two species were differentiated mainly by crest size a ...
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Nodosaurid
Nodosauridae is a family of ankylosaurian dinosaurs, from the Late Jurassic to the Late Cretaceous period in what is now North America, South America, Europe, and Asia. Description Nodosaurids, like their close relatives the ankylosaurids, were heavily armored dinosaurs adorned with rows of bony armor nodules and spines (osteoderms), which were covered in keratin sheaths. All nodosaurids, like other ankylosaurians, were medium-sized to large, heavily built, quadrupedal, herbivorous dinosaurs, possessing small, leaf-shaped teeth. Unlike ankylosaurids, nodosaurids lacked mace-like tail clubs, instead having flexible tail tips. Many nodosaurids had spikes projecting outward from their shoulders. One particularly well-preserved nodosaurid "mummy", known as the Suncor nodosaur (''Borealopelta markmitchelli''), preserved a nearly complete set of armor in life position, as well as the keratin covering and mineralized remains of the underlying skin, which indicate reddish dorsal pigmen ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily "typ ...
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Braincase
In human anatomy, the neurocranium, also known as the braincase, brainpan, or brain-pan is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calvaria or skullcap. The remainder of the skull is the facial skeleton. In comparative anatomy, neurocranium is sometimes used synonymously with endocranium or chondrocranium. Structure The neurocranium is divided into two portions: * the membranous part, consisting of flat bones, which surround the brain; and * the cartilaginous part, or chondrocranium, which forms bones of the base of the skull. In humans, the neurocranium is usually considered to include the following eight bones: * 1 ethmoid bone * 1 frontal bone * 1 occipital bone * 2 parietal bones * 1 sphenoid bone * 2 temporal bones The ossicles (three on each side) are usually not included as bones of the neurocranium. There may variably also be extra sutural bones present. Below the ne ...
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Theropod
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally carnivorous, although a number of theropod groups evolved to become herbivores and omnivores. Theropods first appeared during the Carnian age of the late Triassic period 231.4 million years ago ( Ma) and included all the large terrestrial carnivores from the Early Jurassic until at least the close of the Cretaceous, about 66 Ma. In the Jurassic, birds evolved from small specialized coelurosaurian theropods, and are today represented by about 10,500 living species. Biology Diet and teeth Theropods exhibit a wide range of diets, from insectivores to herbivores and carnivores. Strict carnivory has always been considered the ancestral diet for theropods as a group, and a wider variety of diets was historically considered a characteri ...
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Crocodilian
Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living relatives of birds, as the two groups are the only known survivors of the Archosauria. Members of the order's total group, the clade Pseudosuchia, appeared about 250 million years ago in the Early Triassic period, and diversified during the Mesozoic era. The order Crocodilia includes the true crocodiles (family Crocodylidae), the alligators and caimans (family Alligatoridae), and the gharial and false gharial (family Gavialidae). Although the term 'crocodiles' is sometimes used to refer to all of these, crocodilians is a less ambiguous vernacular term for members of this group. Large, solidly built, lizard-like reptiles, crocodilians have long flattened snouts, laterally compressed tails, and eyes, ears, and nostrils at the top o ...
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Turtle
Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked turtles), which differ in the way the head retracts. There are 360 living and recently extinct species of turtles, including land-dwelling tortoises and freshwater terrapins. They are found on most continents, some islands and, in the case of sea turtles, much of the ocean. Like other amniotes (reptiles, birds, and mammals) they breathe air and do not lay eggs underwater, although many species live in or around water. Turtle shells are made mostly of bone; the upper part is the domed carapace, while the underside is the flatter plastron or belly-plate. Its outer surface is covered in scales made of keratin, the material of hair, horns, and claws. The carapace bones develop from ribs that grow sideways and develop into broad flat plates th ...
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