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Jungle Toad
''Barbourula'' is a genus of amphibian commonly referred to as jungle toads. They are small toads of the fire-bellied toad family, Bombinatoridae, found in the Philippines and Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and ea .... Species References Amphibian Species of the World, ''Barbourula''Living Underworld, Bombinatoridae
Amphibian genera
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Barbourula Busuangensis
''Barbourula busuangensis'' is a species of frog in the family Bombinatoridae. It is also known under the common names Philippine flat-headed frog, Palawan flat-headed frog, Busuanga jungle toad, Busuanga disk-tongued toad, and Philippine aquatic frog. It is endemic to the Busuanga, Culion, Balabac, and Palawan islands in the Philippines. It is an inhabitant of clear-water streams and is threatened by habitat loss. Description ''Barbourula busuangensis'' is a large aquatic species of frog. Its body is flattened dorso-ventrally and its nostrils and eyes are located at the top of its head. It has no external tympani. Both fore and hind limbs are robust and powerful and the digits of the hands are webbed as well as those of the feet. The surface of the body is covered with fine tubercles. This frog is a dark greenish-black in colour with some faint pale green hatching and spotting on the hind limbs. Distribution and habitat This frog is endemic to the western Philippines where it i ...
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Edward Harrison Taylor
Edward Harrison Taylor (April 23, 1889 – June 16, 1978) was an American herpetologist from Missouri. Family Taylor was born in Maysville, Missouri, to George and Loretta Taylor. He had an older brother, Eugene. Education Taylor studied at the University of Kansas in Lawrence, Kansas, graduating with a B.A. in 1912. Field trips during his time at the University of Kansas with Dr. Clarence McClung and Dr. Roy Moody helped prepare Taylor for his future endeavors. Between 1916 and 1920 he returned briefly to Kansas to finish his M.A. Career Upon completing his bachelor's degree, Taylor went to the Philippines, where at first he held a teacher's post in a village in central Mindanao. He collected and studied the local herpetofauna extensively and published many papers. He returned to the Philippines after completing his master's degree and was appointed Chief of Fisheries in Manila. On his many survey trips he continued collecting and studying fishes and reptiles of the islan ...
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Gladwyn Kingsley Noble
Gladwyn Kingsley Noble (September 20, 1894 – December 9, 1940) was an American zoologist who served as the head curator for the Department of Herpetology and the Department of Experimental Biology at the American Museum of Natural History. Noble received bachelor's and master's degrees from Harvard University in 1917 and 1918, respectively, and a Ph.D. from Columbia University in 1922. He joined the herpetology department in 1922 as a research assistant and assistant curator in 1917, and became the chairman of the department in 1924. He later formed the Department of Experimental Biology in 1928, and served as the chairman of both departments until his death on December 9, 1940, from a streptococcal throat infection. Noble is the taxon author of 20 new species of reptiles. A species of lizard, ''List of Anolis lizards, Anolis noblei'', is named in his honor. Also, a subspecies of lizard, ''Sphaerodactylus darlingtoni, Sphaerodactylus darlingtoni noblei'', is named in his honor. ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Amphibian
Amphibians are tetrapod, four-limbed and ectothermic vertebrates of the Class (biology), class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial animal, terrestrial, fossorial, arboreal or freshwater aquatic ecosystems. Thus amphibians typically start out as larvae living in water, but some species have developed behavioural adaptations to bypass this. The young generally undergo metamorphosis from larva with gills to an adult air-breathing form with lungs. Amphibians use their skin as a secondary respiratory surface and some small terrestrial salamanders and frogs lack lungs and rely entirely on their skin. They are superficially similar to reptiles like lizards but, along with mammals and birds, reptiles are amniotes and do not require water bodies in which to breed. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators; in re ...
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Bombinatoridae
Bombinatoridae is a family of toads found in Eurasia. Species of the family have flattened bodies and some are highly toxic. Taxonomy and systematics Fossil specimens of the genus ''Bombina'' are known from the Pliocene to the Pleistocene. The earliest fossil specimens are '' Eobarbourula'' from the Eocene of India, and '' Hatzegobatrachus'' from Late Cretaceous of Hateg island, Romania. The genus ''Barbourula'' was considered to be situated intermediate between ''Discoglossus'' and ''Bombina'', but closer to the latter, so was added to the Bombinatoridae when that family was split from the Discoglossidae. Genera Currently, there are two extant and at least two extinct genera recognised in the family Bombinatoridae:Bombinatoridae
at the

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Philippines
The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republika sang Filipinas * ibg, Republika nat Filipinas * ilo, Republika ti Filipinas * ivv, Republika nu Filipinas * pam, Republika ning Filipinas * krj, Republika kang Pilipinas * mdh, Republika nu Pilipinas * mrw, Republika a Pilipinas * pag, Republika na Filipinas * xsb, Republika nin Pilipinas * sgd, Republika nan Pilipinas * tgl, Republika ng Pilipinas * tsg, Republika sin Pilipinas * war, Republika han Pilipinas * yka, Republika si Pilipinas In the recognized optional languages of the Philippines: * es, República de las Filipinas * ar, جمهورية الفلبين, Jumhūriyyat al-Filibbīn is an archipelagic country in Southeast Asia. It is situated in the western Pacific Ocean and consists of around 7,641 islands t ...
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Borneo
Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and east of Sumatra. The island is politically divided among three countries: Malaysia and Brunei in the north, and Indonesia to the south. Approximately 73% of the island is Indonesian territory. In the north, the East Malaysian states of Sabah and Sarawak make up about 26% of the island. The population in Borneo is 23,053,723 (2020 national censuses). Additionally, the Malaysian federal territory of Labuan is situated on a small island just off the coast of Borneo. The sovereign state of Brunei, located on the north coast, comprises about 1% of Borneo's land area. A little more than half of the island is in the Northern Hemisphere, including Brunei and the Malaysian portion, while the Indonesian portion spans the Northern and Southern hemisph ...
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Barbourula Busuangensis05
''Barbourula'' is a genus of amphibian commonly referred to as jungle toads. They are small toads of the fire-bellied toad family, Bombinatoridae, found in the Philippines and Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and ea .... Species References Amphibian Species of the World, ''Barbourula''Living Underworld, Bombinatoridae
Amphibian genera
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Philippine Flat-headed Frog
''Barbourula busuangensis'' is a species of frog in the family Bombinatoridae. It is also known under the common names Philippine flat-headed frog, Palawan flat-headed frog, Busuanga jungle toad, Busuanga disk-tongued toad, and Philippine aquatic frog. It is endemic to the Busuanga, Culion, Balabac, and Palawan islands in the Philippines. It is an inhabitant of clear-water streams and is threatened by habitat loss. Description ''Barbourula busuangensis'' is a large aquatic species of frog. Its body is flattened dorso-ventrally and its nostrils and eyes are located at the top of its head. It has no external tympani. Both fore and hind limbs are robust and powerful and the digits of the hands are webbed as well as those of the feet. The surface of the body is covered with fine tubercles. This frog is a dark greenish-black in colour with some faint pale green hatching and spotting on the hind limbs. Distribution and habitat This frog is endemic to the western Philippines where it i ...
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Bornean Flat-headed Frog
The Bornean flat-headed frog (''Barbourula kalimantanensis'') is a species of frog in the family Bombinatoridae. Although many salamanders are lungless, the Bornean flat-headed frog is the first frog known to have no lungs. Djoko Iskandar, an Indonesian zoologist, first described the Bornean flat-headed frog based on a single specimen. However, he did not know the frog was lungless until he was in the field with David Bickford's expedition. Bickford ''et al.'' were able to dissect some of the specimens and discovered for the first time that the frogs had no lungs. The frog breathes entirely through its skin, and its internal organs (the stomach, spleen and the liver) take up much of the space which normally would be filled by the lungs. By retaining the lunglessness of their tadpole stage, the frogs are much flatter than typical frogs, which might help absorb oxygen and avoid being swept away in fast streams. Description This frog grows to a snout-to-vent length of for females a ...
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Barbourula
''Barbourula'' is a genus of amphibian commonly referred to as jungle toads. They are small toads of the fire-bellied toad family, Bombinatoridae, found in the Philippines and Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and ea .... Species References Amphibian Species of the World, ''Barbourula''Living Underworld, Bombinatoridae
Amphibian genera
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