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Ikanogavialis
''Ikanogavialis'' is an extinct genus of gavialid crocodilian. Fossils have been found in the Urumaco Formation in Urumaco, Venezuela and the Solimões Formation of Brazil. The strata from which remains are found are late Miocene in age, rather than Pliocene as was once thought. A possible member of this genus survived into the Late Holocene on Muyua or Woodlark Island in Papua New Guinea. Description ''Ikanogavialis'' had a dorsoventrally deep snout and a distinctive notch between the dentary and maxillary alveoli. The external nares projected anterodorsally from the rostrum. This can be seen as a plesiomorphic characteristic in crocodilians, but given that the earliest gavialoids possessed dorsally projecting external nares, this feature can be seen as having been a reversal from the gavialoid apomorphy back to the crocodilian plesiomorphy rather than having been directly obtained from an early crocodilian ancestor. Species The type species of ''Ikanogavialis'' is ''I. ga ...
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Piscogavialis
''Piscogavialis'' is an extinct monospecific genus of gryposuchine gavialid crocodylian. The only species yet known is ''P. jugaliperforatus''. Fossils of ''Piscogavialis'' have been found from the Mio-Pliocene Pisco Formation of the Sacaco Basin in southern Peru in 1998,''Piscogavialis''
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where it coexisted with the much smaller gavialid '' Sacacosuchus''. ''Piscogavialis'' is known only from a single specimen, but it represents some of the best preserved gavialid material known from

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Gryposuchus
''Gryposuchus'' is an extinct genus of gavialid crocodilian. Fossils have been found from Argentina, Colombia, Venezuela, Brazil and the Peruvian Amazon. The genus existed during the Miocene epoch (Colhuehuapian to Huayquerian). One recently described species, ''G. croizati'', grew to an estimated length of . ''Gryposuchus'' is the type genus of the subfamily Gryposuchinae, although a 2018 study indicates that Gryposuchinae and ''Gryposuchus'' might be paraphyletic and rather an evolutionary grade towards the gharial. Species The type species of ''Gryposuchus'' is ''G. jessei'', named based on a well-preserved rostrum collected along the Pauini River of Brazil in 1912. The specimen was probably destroyed during World War II by the 1943 bombing of Hamburg. Another specimen named UFAC 1272, consisting of a premaxilla and maxilla, was discovered in the nearby Sena Madureia locality of the late Miocene Solimões Formation, in and referred to the species in 1997. ''G. jesse ...
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Urumaco Formation
The Urumaco Formation is a formation in Venezuela that includes deposits from the Late Miocene. It is the site of several "giant forms": the turtles, crocodiles, sloths and rodents of Urumaco are among the largest of their groups. Location The Urumaco formation is located in the Urumaco region in the Caribbean coastal Falcón state. The deposits date from 10 to 5.3 million years ago and the Urumaco formation was deposited in an area with large rivers, swamps, estuaries, lagoons and shallow coastal seas. These conditions in the Late Miocene contrast strongly with the current dry environment in the area today. Fauna Cartilaginous fish There are 21 known species of cartilaginous fishes from the Urumaco Formation, belonging to the orders Lamniformes, Carcharhiniformes, Myliobatiformes and Rajiformes. ''Carcharhinus caquetius'' is an endemic species of predator shark from Urumaco. A large number of well-preserved fossils of the sawfish ''Pristis rostra'' have been found in th ...
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Urumaco Formation
The Urumaco Formation is a formation in Venezuela that includes deposits from the Late Miocene. It is the site of several "giant forms": the turtles, crocodiles, sloths and rodents of Urumaco are among the largest of their groups. Location The Urumaco formation is located in the Urumaco region in the Caribbean coastal Falcón state. The deposits date from 10 to 5.3 million years ago and the Urumaco formation was deposited in an area with large rivers, swamps, estuaries, lagoons and shallow coastal seas. These conditions in the Late Miocene contrast strongly with the current dry environment in the area today. Fauna Cartilaginous fish There are 21 known species of cartilaginous fishes from the Urumaco Formation, belonging to the orders Lamniformes, Carcharhiniformes, Myliobatiformes and Rajiformes. ''Carcharhinus caquetius'' is an endemic species of predator shark from Urumaco. A large number of well-preserved fossils of the sawfish ''Pristis rostra'' have been found in th ...
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Gavialid
Gavialidae is a family of large semiaquatic crocodilians with elongated, narrow snouts. Gavialidae consists of two living species, the gharial (''Gavialis gangeticus'') and the false gharial (''Tomistoma schlegelii''), both occurring in Asia. Many extinct members are known from a broader range, including the recently extinct ''Hanyusuchus''. Gavialids are generally regarded as lacking the jaw strength to capture the large mammalian prey favoured by crocodiles and alligators of similar size so their thin snout is best used to catch fish, however the false gharial has been found to have a generalist diet with mature adults preying upon larger vertebrates, such as ungulates. Taxonomy The family Gavialidae was proposed by Arthur Adams in 1854 for reptiles with a very long and slender muzzle, webbed feet and nearly equal teeth. It is currently recognized as a crown group, meaning that it only includes the last common ancestor of all extant (living) gavialids (the gharial and false gha ...
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Crocodilia
Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living relatives of birds, as the two groups are the only known survivors of the Archosauria. Members of the order's total group, the clade Pseudosuchia, appeared about 250 million years ago in the Early Triassic period, and diversified during the Mesozoic era. The order Crocodilia includes the true crocodiles (family Crocodylidae), the alligators and caimans (family Alligatoridae), and the gharial and false gharial (family Gavialidae). Although the term 'crocodiles' is sometimes used to refer to all of these, crocodilians is a less ambiguous vernacular term for members of this group. Large, solidly built, lizard-like reptiles, crocodilians have long flattened snouts, laterally compressed tails, and eyes, ears, and nostrils at the top ...
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Siquisiquesuchus
''Siquisiquesuchus'' (meaning " Siquisique crocodile" after the town in Lara, Venezuela, near where the first described specimens were found) is an extinct genus of gavialid crocodilian. It is known from cranial remains and a few postcranial bones found in Miocene-age rocks of the Castillo Formation in northwestern Venezuela. Description ''Siquisiquesuchus'' is based on the holotype MBLUZ–P–5050, a nearly complete skull and lower jaws. It was found near Lara in rocks of the Early Miocene-age Castillo Formation. Two other partial skulls, partial vertebrae, a thigh bone, partial upper arm, and partial shin bone were recovered from another locality. These bones were described by Christopher Brochu and Ascanio Rincón in 2004. The type species is ''S. venezuelensis''. Like other gavialoids, ''Siquisiquesuchus'' had a long, narrow rostrum on its skull, accounting for approximately 60% of the skull's length. The number of teeth in the premaxillary bones at the tip of the snou ...
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Woodlark Island
Woodlark Island, known to its inhabitants simply as Woodlark or Muyua, is the main island of the Woodlark Islands archipelago, located in Milne Bay Province and the Solomon Sea, Papua New Guinea. Although no formal census has been conducted since 1990, the current population is estimated at approximately 6000 people (see section below on Population Issues). There is one primary school on the island (in Kulumadau) that teaches about 200 students (60 students are from outer islands); to attend highschool/secondary school, and all children must travel to Alotau on the mainland. Etymology Woodlark Island is also called Woodlark or Woodlarks by English language speakers. It is called Murua by the inhabitants of some other islands in the province. The wider Woodlark Islands group also consists of Madau and Nusam to the west, Nubara to the east, and the Marshall Bennett group to the southwest. History The Australian whaling ship (Captain George Grimes) called in the 1830s and th ...
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Gavialis
''Gavialis'' is a genus of crocodylians that includes the living gharial ''Gavialis gangeticus'' and one known extinct species, '' Gavialis bengawanicus.'' ''G. gangeticus'' comes from the Indian Subcontinent, while ''G. bengawanicus'' is known from Java. ''Gavialis'' likely first appeared in the Indian Subcontinent in the Pliocene and dispersed into the Malay Archipelago through a path called the Siva–Malayan route in the Quaternary. Remains attributed to ''Gavialis'' have also been found on Sulawesi and Woodlark Island east of the Wallace Line, suggesting a prehistoric lineage of ''Gavialis'' was able to traverse marine environments and reach places possibly as far as western Oceania. The genus ''Gavialis'' was reevaluated in 2018 based on specimens in the Natural History Museum, London that were collected in the Sivalik Hills. The author concluded that ''G. gangeticus'' and '' G. bengawanicus'' are the only two species in the genus ''Gavialis'', with '' G. hysudricus'' a ...
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Maxilla
The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxillary bones are fused at the intermaxillary suture, forming the anterior nasal spine. This is similar to the mandible (lower jaw), which is also a fusion of two mandibular bones at the mandibular symphysis. The mandible is the movable part of the jaw. Structure In humans, the maxilla consists of: * The body of the maxilla * Four processes ** the zygomatic process ** the frontal process of maxilla ** the alveolar process ** the palatine process * three surfaces – anterior, posterior, medial * the Infraorbital foramen * the maxillary sinus * the incisive foramen Articulations Each maxilla articulates with nine bones: * two of the cranium: the frontal and ethmoid * seven of the face: the nasal, zygomatic, lacrimal, inferior n ...
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Gavialoids
Gavialoidea is one of three superfamilies of crocodylians, the other two being Alligatoroidea and Crocodyloidea. Although many extinct species are known, only the gharial ''Gavialis gangeticus'' and the false gharial ''Tomistoma schlegelii'' are alive today, with ''Hanyusuchus'' having become extinct in the last few centuries. Extinct South American gavialoids likely dispersed in the mid Tertiary from Africa and Asia. Fossil remains of the Puerto Rican gavialoid '' Aktiogavialis puertorisensis'' were discovered in a cave located in San Sebastián, Puerto Rico and dated to the Oligocene. This individual is thought to have crossed the Atlantic coming from Africa, indicating that this species was able to withstand saltwater. Classification Gavialoidea is cladistically defined as ''Gavialis gangeticus'' (the gharial) and all crocodylians closer to it than to ''Alligator mississippiensis'' (the American alligator) or ''Crocodylus niloticus'' (the Nile crocodile). This is a stem-bas ...
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PeerJ
''PeerJ'' is an open access peer-reviewed scientific mega journal covering research in the biological and medical sciences. It is published by a company of the same name that was co-founded by CEO Jason Hoyt (formerly at Mendeley) and publisher Peter Binfield (formerly at '' PLOS One''), with initial financial backing of US$950,000 from O'Reilly Media's O'Reilly AlphaTech Ventures, and later funding from Sage Publishing. PeerJ officially launched in June 2012, started accepting submissions on December 3, 2012, and published its first articles on February 12, 2013. The company is a member of CrossRef, CLOCKSS, ORCID, and the Open Access Scholarly Publishers Association. The company's offices are in Corte Madera (California, USA), and London (Great Britain). Submitted research is judged solely on scientific and methodological soundness (as at '' PLoS ONE''), with a facility for peer reviews to be published alongside each paper. Business model ''PeerJ'' uses a business model th ...
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