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Hybodonts
Hybodontiformes, commonly called hybodonts, are an extinct group of shark-like chondrichthyans, which existed from the late Devonian to the Late Cretaceous. They form the group of Elasmobranchii closest to neoselachians, the clade of modern sharks and rays. Hybodonts were named and are distinguished based on their conical tooth shape. They are also noted for the presence of a spine on each of their two dorsal fins. They were abundant in marine and freshwater environments during the late Paleozoic and early Mesozoic, but were rare in open marine environments by the end of the Jurassic, having been largely replaced by modern sharks, though they were still common in freshwater and marginal marine habitats. They survived until the end of the Cretaceous, before going extinct. Etymology The term hybodont comes from the Greek word ''ὕβος'' or ''ὑβός'' meaning hump or hump-backed and ''ὀδούς, ὀδοντ'' meaning tooth. This name was given based on their conical ...
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Hybodus Fraasi (fossil)
''Hybodus'' (from el, ύβος , 'crooked' and el, ὀδούς 'tooth') is an extinct genus of hybodont, a group of shark-like elasmobranchs that lived from the Late Devonian to the end of the Cretaceous. Species closely related to the type species ''Hybodus reticulatus'' lived during the Early Jurassic epoch. Numerous species have been assigned to ''Hybodus'' spanning a large period of time, and it is currently considered a wastebasket taxon that is 'broadly polyphyletic' and requires reexamination. The first fossilized teeth from ''Hybodus'' were found in England around 1845; since then teeth (and dorsal spines) have been recovered from Europe. During the Triassic, Jurassic, and Cretaceous periods, the hybodonts were especially successful and could be found in shallow seas around the world. For reasons that are not fully understood, the hybodonts became extinct near the end of the Late Cretaceous period. Description ''Hybodus'' species grew to about in length. It was not ver ...
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Jurassic
The Jurassic ( ) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period of the Mesozoic Era and is named after the Jura Mountains, where limestone strata from the period were first identified. The start of the Jurassic was marked by the major Triassic–Jurassic extinction event, associated with the eruption of the Central Atlantic Magmatic Province. The beginning of the Toarcian Stage started around 183 million years ago and is marked by an extinction event associated with widespread oceanic anoxia, ocean acidification, and elevated temperatures likely caused by the eruption of the Karoo-Ferrar large igneous provinces. The end of the Jurassic, however, has no clear boundary with the Cretaceous and is the only boundary between geological periods to remain formally undefined. By the beginning of the Jurassi ...
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Sharks
Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachimorpha (or Selachii) and are the sister group to the rays. However, the term "shark" has also been used to refer to all extinct members of Chondrichthyes with a shark-like morphology, such as hybodonts and xenacanths. The oldest modern sharks are known from the Early Jurassic. They range in size from the small dwarf lanternshark (''Etmopterus perryi''), a deep sea species that is only in length, to the whale shark (''Rhincodon typus''), the largest fish in the world, which reaches approximately in length. Sharks are found in all seas and are common to depths up to . They generally do not live in freshwater, although there are a few known exceptions, such as the bull shark and the river shark, which can be found in both seawater and fre ...
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Neoselachii
Elasmobranchii () is a subclass of Chondrichthyes or cartilaginous fish, including sharks (superorder Selachii), rays, skates, and sawfish (superorder Batoidea). Members of this subclass are characterised by having five to seven pairs of gill clefts opening individually to the exterior, rigid dorsal fins and small placoid scales on the skin. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. The details of this jaw anatomy vary between species, and help distinguish the different elasmobranch clades. The pelvic fins in males are modified to create claspers for the transfer of sperm. There is no swim bladder; instead, these fish maintain buoyancy with large livers rich in oil. The definition of the clade is unclear with respect to fossil chondrichthyans. It has been used by different authors as equivalent to Neoselachii (the clade including modern sharks and rays and their last common ancestor) or for a ...
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Asteracanthus
''Asteracanthus'' (from el, ἀστήρ , 'star' and el, ἄκανθα , 'spine') is an extinct genus of hybodontiform, known from the Middle Jurassic (Bathonian) to the Early Cretaceous (Valanginian). Description Astercanthus was among the largest known hybodontiformes, reaching a length of 2-3 metres. The dentition of ''Astercanthus'' is high crowned and multicusped. Fossil records This genus has been reported from the Middle Triassic to the Cretaceous, though the genus as currently circumscribed dates from the Bathonian-Valanginian, predominantly of Europe. Fossils are found in the marine strata of United States, Iran, Switzerland, Madagascar, Morocco and Europe. A complete skeleton was described in 2021 from the Late Jurassic (Tithonian) aged Solnhofen Limestone. Previously considered synonymous, the genus '' Strophodus'' (Middle Triassic-Late Cretaceous) is now considered distinct, with the teeth of ''Asteracanthus'' having more in common with '' Hybodus'' and '' Egert ...
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Strophodus
''Strophodus'' is an extinct genus of hybodonts known from the Triassic to Cretaceous. It has heavily rounded, durophagous teeth. It has long been confused with ''Asteracanthus'' due to the fin spines of the latter being found associated with the teeth of ''Strophodus.'' However, both genera can now be reliably be distinguished base on the morphology of both the fin spines and teeth. Species After * ''S. reticulatus'' Agassiz, 1838 Middle-Late Jurassic (Bathonian–Tithonian) England, France, Germany, Hungary and Switzerland. A similar form ''S. cf. reticulatus'' is known from the Middle Triassic of Switzerland * ''S. smithwoodwardi'' (Peyer, 1946) Early Jurassic (Toarcian) Switzerland * ''S. dunaii'' (Szabó & Főzy (2020) Middle Jurassic (Aalenian), Hungary * ''S. tenuis'' Agassiz, 1838 Middle Jurassic (Aalenian-Bathonian) Germany, England * ''S. longidens'' Agassiz, 1838 Middle Jurassic (Bathonian) France * ''S. magnus'' Agassiz, 1838 Middle Jurassic (Bathonian) France, Ind ...
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Elasmobranchii
Elasmobranchii () is a subclass of Chondrichthyes or cartilaginous fish, including sharks (superorder Selachii), rays, skates, and sawfish (superorder Batoidea). Members of this subclass are characterised by having five to seven pairs of gill clefts opening individually to the exterior, rigid dorsal fins and small placoid scales on the skin. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. The details of this jaw anatomy vary between species, and help distinguish the different elasmobranch clades. The pelvic fins in males are modified to create claspers for the transfer of sperm. There is no swim bladder; instead, these fish maintain buoyancy with large livers rich in oil. The definition of the clade is unclear with respect to fossil chondrichthyans. It has been used by different authors as equivalent to Neoselachii (the clade including modern sharks and rays and their last common ancestor) or f ...
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Elasmobranch
Elasmobranchii () is a subclass of Chondrichthyes or cartilaginous fish, including sharks (superorder Selachii), rays, skates, and sawfish (superorder Batoidea). Members of this subclass are characterised by having five to seven pairs of gill clefts opening individually to the exterior, rigid dorsal fins and small placoid scales on the skin. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. The details of this jaw anatomy vary between species, and help distinguish the different elasmobranch clades. The pelvic fins in males are modified to create claspers for the transfer of sperm. There is no swim bladder; instead, these fish maintain buoyancy with large livers rich in oil. The definition of the clade is unclear with respect to fossil chondrichthyans. It has been used by different authors as equivalent to Neoselachii (the clade including modern sharks and rays and their last common ancestor) or fo ...
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Chondrichthyes
Chondrichthyes (; ) is a class that contains the cartilaginous fishes that have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or ''bony fishes'', which have skeletons primarily composed of bone tissue. Chondrichthyes are jawed vertebrates with paired fins, paired nares, scales, and a heart with its chambers in series. Extant chondrichthyes range in size from the 10 cm (3.9 in) finless sleeper ray to the 10 m (32 ft) whale shark. The class is divided into two subclasses: Elasmobranchii ( sharks, rays, skates, and sawfish) and Holocephali (chimaeras, sometimes called ghost sharks, which are sometimes separated into their own class). Within the infraphylum Gnathostomata, cartilaginous fishes are distinct from all other jawed vertebrates. Anatomy Skeleton The skeleton is cartilaginous. The notochord is gradually replaced by a vertebral column during development, except in Holocephali, where the notochord stays intact. In some d ...
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Robustness (morphology)
In biology, robustness is used to describe a taxon with a stronger and heavier build ( morphology) when compared to a related gracile taxon. The terms are used in contrast to one another. The term is used by physical anthropologists and paleoanthropologists to refer to a big-boned and muscular body. For example, members of the genus '' Sapajus'' have robust body types and are called the robust capuchin monkeys while members of the genus '' Cebus'' have gracile body types and are called the gracile capuchin monkeys. Male and female members of the same species may display sexual dimorphism and have robust and gracile morphologies. The terms "robust" vs. "gracile" are used in the context of human evolution, to distinguish: *"robust" vs. "gracile" australopithecines, see '' Paranthropus'' *"robust" archaic humans vs. "gracile" anatomically modern humans *"robust" early modern humans (Cro-Magnon) vs. "gracile" Epipaleolithic humans (Combe-Capelle Combe-Capelle is a Paleolithic ...
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Cartilaginous
Cartilage is a resilient and smooth type of connective tissue. In tetrapods, it covers and protects the ends of long bones at the joints as articular cartilage, and is a structural component of many body parts including the rib cage, the neck and the bronchial tubes, and the intervertebral discs. In other taxa, such as chondrichthyans, but also in cyclostomes, it may constitute a much greater proportion of the skeleton. It is not as hard and rigid as bone, but it is much stiffer and much less flexible than muscle. The matrix of cartilage is made up of glycosaminoglycans, proteoglycans, collagen fibers and, sometimes, elastin. Because of its rigidity, cartilage often serves the purpose of holding tubes open in the body. Examples include the rings of the trachea, such as the cricoid cartilage and carina. Cartilage is composed of specialized cells called chondrocytes that produce a large amount of collagenous extracellular matrix, abundant ground substance that is rich in pr ...
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Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began dominat ...
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