Hulsanpes
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Hulsanpes
''Hulsanpes'' ( meaning " Khulsan foot") is a genus of halszkaraptorine theropod dinosaurs that lived during the Late Cretaceous in what is now the Barun Goyot Formation of Mongolia, about 75-72 million years ago. The remains were found in 1970 and formally described in 1982 by Halszka Osmólska, who noted that the genus is represented by an immature individual. ''Hulsanpes'' represents the first record of the basal dromaeosaurid subfamily Halszkaraptorinae. History of discovery The first fossil remains of ''Hulsanpes'' were discovered in 1970 during the third Polish-Mongolian expedition at Khulsan in the Gobi Desert. Twelve years later, the type species, ''Hulsanpes perlei'', was named and described by the Polish palaeontologist Halszka Osmólska in 1982. ''Hulsanpes'' is based on the holotype fossil specimen ZPAL MgD-I/173, uncovered in a sandstone layer of the Barun Goyot Formation, dating from the Late Campanian (roughly 72 million years ago). It consists of a partial brain ...
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Hulsanpes Braincase Wall
''Hulsanpes'' ( meaning " Khulsan foot") is a genus of halszkaraptorine theropod dinosaurs that lived during the Late Cretaceous in what is now the Barun Goyot Formation of Mongolia, about 75-72 million years ago. The remains were found in 1970 and formally described in 1982 by Halszka Osmólska, who noted that the genus is represented by an immature individual. ''Hulsanpes'' represents the first record of the basal dromaeosaurid subfamily Halszkaraptorinae. History of discovery The first fossil remains of ''Hulsanpes'' were discovered in 1970 during the third Polish-Mongolian expedition at Khulsan in the Gobi Desert. Twelve years later, the type species, ''Hulsanpes perlei'', was named and described by the Polish palaeontologist Halszka Osmólska in 1982. ''Hulsanpes'' is based on the holotype fossil specimen ZPAL MgD-I/173, uncovered in a sandstone layer of the Barun Goyot Formation, dating from the Late Campanian (roughly 72 million years ago). It consists of a partial brainc ...
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Halszkaraptorine
Halszkaraptorinae is a basal ("primitive") subfamily of Dromaeosauridae (or possibly Unenlagiidae) that includes the enigmatic genera ''Halszkaraptor'', ''Natovenator'', ''Mahakala'', and ''Hulsanpes''. Halszkaraptorines are definitively known only from Late Cretaceous strata in Asia, specifically in Mongolia. Following the recent discovery of ''Natovenator'', a member of the subfamily, the group is confirmed to have a semiaquatic lifestyle. History of discovery The first known remains of halszkaraptorines were found in sandstone sediments in 1970 during a Polish-Mongolian expedition at the Barun Goyot Formation, Gobi Desert. Later in 1982 they were described by Polish palaeontologist Halszka Osmólska and used as the holotype for the new genus and species '' Hulsanpes perlei'', honoring the Mongolian paleontologist Altangerel Perle. Although the affinities of these fossils were not fully understood, they were tentatively interpreted as belonging to Deinonychosauria. In 1992 dro ...
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Halszkaraptorinae
Halszkaraptorinae is a basal ("primitive") subfamily of Dromaeosauridae (or possibly Unenlagiidae) that includes the enigmatic genera ''Halszkaraptor'', ''Natovenator'', ''Mahakala'', and ''Hulsanpes''. Halszkaraptorines are definitively known only from Late Cretaceous strata in Asia, specifically in Mongolia. Following the recent discovery of ''Natovenator'', a member of the subfamily, the group is confirmed to have a semiaquatic lifestyle. History of discovery The first known remains of halszkaraptorines were found in sandstone sediments in 1970 during a Polish-Mongolian expedition at the Barun Goyot Formation, Gobi Desert. Later in 1982 they were described by Polish palaeontologist Halszka Osmólska and used as the holotype for the new genus and species '' Hulsanpes perlei'', honoring the Mongolian paleontologist Altangerel Perle. Although the affinities of these fossils were not fully understood, they were tentatively interpreted as belonging to Deinonychosauria. In 1992 dro ...
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Dromaeosauridae
Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning 'runner', and ('), meaning 'lizard'. In informal usage, they are often called raptors (after ''Velociraptor''), a term popularized by the film '' Jurassic Park''; a few types include the term "raptor" directly in their name and have come to emphasize their bird-like appearance and speculated bird-like behavior. Dromaeosaurid fossils have been found across the globe in North America, Europe, Africa, Asia, South America and Antarctica, with some fossils giving credence to the possibility that they inhabited Australia as well. They first appeared in the mid-Jurassic Period (late Bathonian stage, about 167 million years ago) and survived until the end of the Cretaceous (Maastrichtian stage, 66 ma), existing until the Cretaceous–Paleogene ex ...
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1982 In Paleontology
Plants Pteridophytes Angiosperms Fish Ray-finned fish Archosauromorphs Newly named dinosaurs Data courtesy of George Olshevsky's dinosaur genera list. Birds Some remarks on newly named birds * ''Hulsanpes perlei'' Osmolska, 1982 was described as a Troodontidae, it is sometimes seen as an Avialae because of having a ginglymoid metatarsal II. Newly named birds Plesiosaurs New taxa Pterosaurs New taxa Crocodylomorpha New taxa References {{reflist, 30em Paleontology Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ... Paleontology 2 ...
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Barun Goyot Formation
The Barun Goyot Formation (also known as Baruungoyot Formation or West Goyot Formation) is a geological formation dating to the Late Cretaceous Period. It is located within and is widely represented in the Gobi Desert Basin, in the Ömnögovi Province of Mongolia. Description It was previously known as the Lower Nemegt Beds occurring beneath the Nemegt Formation and above the Djadokhta Formation. It has been suggested that the Djadokhta and Barun Goyot Formations are lower and upper parts, respectively, of the same lithological unit and the boundary between the two does not exist. The stratotype of the Barun Goyot Formation is the Khulsan locality, east of Nemegt. At Nemegt, only the uppermost barungoyotian beds are visible. The ''Red Beds of Khermeen Tsav'' are also considered part of the Barun Goyot Formation. It is approximately in thickness,Gradzinski, R.; & Jerzykiewicz, T. (1974). Sedimentation of the Barun Goyot formation. Palaeontologica Polonica, 30, 111-146. and was l ...
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Halszka Osmólska
Halszka Osmólska (September 15, 1930 – March 31, 2008) was a Polish paleontologist who had specialized in Mongolian dinosaurs. Biography She was born in 1930 in Poznań. In 1949, she began to study biology at Faculty of Biology and Earth Sciences of the University of Poznań before moving to Warsaw and studying at the Warsaw University, which she graduated from in 1955. Since then she worked at the Institute of Paleobiology of the Polish Academy of Sciences (PAN). Between 1983–1988, she served as the institute's director. She was a member of the Polish–Mongolian expeditions to the Gobi Desert (1963–1965 and 1967–1971) and she described many finds from these rocks, often with Teresa Maryańska. Among the dinosaurs she described are: * ''Elmisaurus'' (and Elmisauridae) (1981), ''Hulsanpes'' (1982), '' Borogovia'' (1987), and ''Bagaraatan'' (1996) * with Maryańska, ''Homalocephale'', ''Prenocephale'', and ''Tylocephale'' (and Pachycephalosauria)(1974), ''Bagaceratops'' ...
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Campanian
The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian spans the time from 83.6 (± 0.2) to 72.1 (± 0.2) million years ago. It is preceded by the Santonian and it is followed by the Maastrichtian. The Campanian was an age when a worldwide sea level rise covered many coastal areas. The morphology of some of these areas has been preserved: it is an unconformity beneath a cover of marine sedimentary rocks. Etymology The Campanian was introduced in scientific literature by Henri Coquand in 1857. It is named after the French village of Champagne in the department of Charente-Maritime. The original type locality was a series of outcrop near the village of Aubeterre-sur-Dronne in the same region. Definition The base of the Campanian Stage is defined as a place in the stratigraphic column wher ...
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Million Years Ago
The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). Together they make a reference system, one to a quantity, the other to a particular place in a year numbering system that is ''time before the present''. Myr is deprecated in geology, but in astronomy ''Myr'' is standard. Where "myr" ''is'' seen in geology it is usually "Myr" (a unit of mega-years). In astronomy it is usually "Myr" (Million years). Debate In geology a debate remains open concerning the use of ''Myr'' (duration) plus ''Ma'' (million years ago) versus using only the term ''Ma''. In either case the term '' Ma'' is used in geology literature conforming to ISO 31-1 (now ISO 80000-3) and NIST 811 recommended practices. Traditional style geology literature is written The "ago" is implied, so that any such year number "X Ma" ...
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Braincase
In human anatomy, the neurocranium, also known as the braincase, brainpan, or brain-pan is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calvaria or skullcap. The remainder of the skull is the facial skeleton. In comparative anatomy, neurocranium is sometimes used synonymously with endocranium or chondrocranium. Structure The neurocranium is divided into two portions: * the membranous part, consisting of flat bones, which surround the brain; and * the cartilaginous part, or chondrocranium, which forms bones of the base of the skull. In humans, the neurocranium is usually considered to include the following eight bones: * 1 ethmoid bone * 1 frontal bone * 1 occipital bone * 2 parietal bones * 1 sphenoid bone * 2 temporal bones The ossicles (three on each side) are usually not included as bones of the neurocranium. There may variably also be extra sutural bones present. Below the ne ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Ant ...
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Metatarsus
The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the medial side (the side of the great toe): the first, second, third, fourth, and fifth metatarsal (often depicted with Roman numerals). The metatarsals are analogous to the metacarpal bones of the hand. The lengths of the metatarsal bones in humans are, in descending order, second, third, fourth, fifth, and first. Structure The five metatarsals are dorsal convex long bones consisting of a shaft or body, a base (proximally), and a head (distally).Platzer 2004, p. 220 The body is prismoid in form, tapers gradually from the tarsal to the phalangeal extremity, and is curved longitudinally, so as to be concave below, slightly convex above. The base or posterior extremity is wedge-shaped, articulating proximally with the tarsal bones, and by it ...
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