Helicoprion
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Helicoprion
''Helicoprion'' is an extinct genus of shark-like eugeneodont fish. Almost all fossil specimens are of spirally arranged clusters of the individuals' teeth, called "tooth whorls", which in life were embedded in the lower jaw. As with most extinct cartilaginous fish, the skeleton is mostly unknown. Fossils of ''Helicoprion'' are known from a 20 million year timespan during the Permian period from the Artinskian stage of the Cisuralian (Early Permian) to the Roadian stage of the Guadalupian (Middle Permian). The closest living relatives of ''Helicoprion'' (and other eugeneodonts) are the chimaeras, though their relationship is very distant. The unusual tooth arrangement is thought to have been an adaption for feeding on soft bodied prey, and may have functioned as a deshelling mechanism for hard bodied cephalopods such as nautiloids and ammonoids. In 2013, systematic revision of ''Helicoprion'' via morphometric analysis of the tooth whorls found only ''H. davisii, H. bessonowi'' ...
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Eugeneodontida
The Eugeneodontida is an extinct and poorly known order of Chondrichthyes, cartilaginous fishes. They possessed "tooth-whorls" on the symphysis of either the lower or both jaws and pectoral fins supported by long radials. They probably lacked pelvic fins and anal fins. The palatoquadrate was either fused to the skull or reduced. Now determined to be within the Holocephali, their closest living relatives are Chimaeridae, ratfish. The eugeneodonts are named after paleontologist Eugene S. Richardson, Jr. The Eugeneodontida disappeared in the Early Triassic. Members of the Eugeneodontida are further classified into different families, the most well-preserved members that have been discovered are commonly placed within the families Helicoprionidae ("spiral saws"), and Edestidae ("those which devour"), the former containing the genera ''Helicoprion'', ''Sarcoprion'', and ''Parahelicoprion'', and the latter containing the genera ''Edestus'', ''Lestrodus'', and ''Metaxyacanthus''. All eu ...
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Cisuralian
The Cisuralian is the first series/epoch of the Permian. The Cisuralian was preceded by the Pennsylvanian and followed by the Guadalupian. The Cisuralian Epoch is named after the western slopes of the Ural Mountains in Russia and Kazakhstan and dates between 298.9 ± 0.15 – 272.3 ± 0.5 Mya. The Cisuralian is often synonymous with the informal terms early Permian or lower Permian. It corresponds approximately with the Wolfcampian in southwestern North America. The series saw the appearance of beetles and flies and was a relatively stable warming period of about 21 million years. Name and background The Cisuralian is the first series or epoch of the Permian. The Cisuralian was preceded by the last Pennsylvanian epoch (Gzhelian) and is followed by the Permian Guadalupian Epoch. The name "Cisuralian" was proposed in 1982, and approved by the International Subcommission on Permian Stratigraphy in 1996. The Cisuralian Epoch is named after the western slopes of the Ural Mountain ...
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Phosphoria Formation
The Phosphoria Formation of the western United States is a geological formation of Early Permian age.Behnken, F.H., Wardlaw, B.R. and Stout, L.N., 1986, Conodont biostratigraphy of the Permian Meade Peak Phosphatic Shale Member, Phosphoria Formation, southeastern Idaho. University of Wyoming Contributions to Geology, v. 24, no. 2, p. 169-190. It represents some 15 million years of sedimentation, reaches a thickness of and covers an area of .Blatt, Harvey and Robert J. Tracy, ''Petrology'', Freeman, 1996, 2nd ed. pp. 345-349 The Phosphoria includes phosphorite beds that are an important source of phosphorus. Many of its shales are rich in organic matter and are petroleum source rocks, and some of its dolomites include petroleum reservoirs. Environment of deposition The Phosphoria Formation was deposited under marine conditions in a foreland basin located between the Paleozoic continental margin and the North American cratonic shelf.Maughan, E.K. 1984. Geological setting and s ...
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Chondrichthyes
Chondrichthyes (; ) is a class that contains the cartilaginous fishes that have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or ''bony fishes'', which have skeletons primarily composed of bone tissue. Chondrichthyes are jawed vertebrates with paired fins, paired nares, scales, and a heart with its chambers in series. Extant chondrichthyes range in size from the 10 cm (3.9 in) finless sleeper ray to the 10 m (32 ft) whale shark. The class is divided into two subclasses: Elasmobranchii (sharks, rays, skates, and sawfish) and Holocephali ( chimaeras, sometimes called ghost sharks, which are sometimes separated into their own class). Within the infraphylum Gnathostomata, cartilaginous fishes are distinct from all other jawed vertebrates. Anatomy Skeleton The skeleton is cartilaginous. The notochord is gradually replaced by a vertebral column during development, except in Holocephali, where the notochord stays intact. In ...
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Permian
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids ( reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of th ...
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Japan
Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north toward the East China Sea, Philippine Sea, and Taiwan in the south. Japan is a part of the Ring of Fire, and spans an archipelago of 6852 islands covering ; the five main islands are Hokkaido, Honshu (the "mainland"), Shikoku, Kyushu, and Okinawa. Tokyo is the nation's capital and largest city, followed by Yokohama, Osaka, Nagoya, Sapporo, Fukuoka, Kobe, and Kyoto. Japan is the eleventh most populous country in the world, as well as one of the most densely populated and urbanized. About three-fourths of the country's terrain is mountainous, concentrating its population of 123.2 million on narrow coastal plains. Japan is divided into 47 administrative prefectures and eight traditional regions. The Greater Tokyo Ar ...
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China
China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and Borders of China, borders fourteen countries by land, the List of countries and territories by land borders, most of any country in the world, tied with Russia. Covering an area of approximately , it is the world's third List of countries and dependencies by area, largest country by total land area. The country consists of 22 provinces of China, provinces, five autonomous regions of China, autonomous regions, four direct-administered municipalities of China, municipalities, and two special administrative regions of China, Special Administrative Regions (Hong Kong and Macau). The national capital is Beijing, and the List of cities in China by population, most populous cit ...
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Kazakhstan
Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eigh ... to Kazakhstan–Russia border, the north and west, China to China–Kazakhstan border, the east, Kyrgyzstan to Kazakhstan–Kyrgyzstan border, the southeast, Uzbekistan to Kazakhstan–Uzbekistan border, the south, and Turkmenistan to Kazakhstan–Turkmenistan border, the southwest, with a coastline along the Caspian Sea. Its capital is Astana, known as Nur-Sultan from 2019 to 2022. Almaty, Kazakhstan's largest city, was the country's capital until 1997. Kazakhstan is the world's largest landlocked country, the largest and northernmost Muslim world, Muslim-majority cou ...
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Norway
Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and the archipelago of Svalbard also form part of Norway. Bouvet Island, located in the Subantarctic, is a dependency of Norway; it also lays claims to the Antarctic territories of Peter I Island and Queen Maud Land. The capital and largest city in Norway is Oslo. Norway has a total area of and had a population of 5,425,270 in January 2022. The country shares a long eastern border with Sweden at a length of . It is bordered by Finland and Russia to the northeast and the Skagerrak strait to the south, on the other side of which are Denmark and the United Kingdom. Norway has an extensive coastline, facing the North Atlantic Ocean and the Barents Sea. The maritime influence dominates Norway's climate, with mild lowland temperatures on the ...
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Laos
Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist state and the only landlocked country in Southeast Asia. At the heart of the Indochinese Peninsula, Laos is bordered by Myanmar and China to the northwest, Vietnam to the east, Cambodia to the southeast, and Thailand to the west and southwest. Its capital and largest city is Vientiane. Present-day Laos traces its historic and cultural identity to Lan Xang, which existed from the 14th century to the 18th century as one of the largest kingdoms in Southeast Asia. Because of its central geographical location in Southeast Asia, the kingdom became a hub for overland trade and became wealthy economically and culturally. After a period of internal conflict, Lan Xang broke into three separate kingdoms: Luang Phrabang, Vientiane and Champa ...
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Russia
Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eighth of Earth's inhabitable landmass. Russia extends across eleven time zones and shares land boundaries with fourteen countries, more than any other country but China. It is the world's ninth-most populous country and Europe's most populous country, with a population of 146 million people. The country's capital and largest city is Moscow, the largest city entirely within Europe. Saint Petersburg is Russia's cultural centre and second-largest city. Other major urban areas include Novosibirsk, Yekaterinburg, Nizhny Novgorod, and Kazan. The East Slavs emerged as a recognisable group in Europe between the 3rd and 8th centuries CE. Kievan Rus' arose as a state in the 9th century, and in 988, it adopted Orthodox Christianity from the ...
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Canada
Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by total area. Its southern and western border with the United States, stretching , is the world's longest binational land border. Canada's capital is Ottawa, and its three largest metropolitan areas are Toronto, Montreal, and Vancouver. Indigenous peoples have continuously inhabited what is now Canada for thousands of years. Beginning in the 16th century, British and French expeditions explored and later settled along the Atlantic coast. As a consequence of various armed conflicts, France ceded nearly all of its colonies in North America in 1763. In 1867, with the union of three British North American colonies through Confederation, Canada was formed as a federal dominion of four provinces. This began an accretion of provinces and ...
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