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Hymenoptera Ecology
Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typically have a special ovipositor for inserting eggs into hosts or places that are otherwise inaccessible. This ovipositor is often modified into a stinger. The young develop through holometabolism (complete metamorphosis)—that is, they have a wormlike larval stage and an inactive pupal stage before they mature. Etymology The name Hymenoptera refers to the wings of the insects, but the original derivation is ambiguous. All references agree that the derivation involves the Ancient Greek πτερόν (''pteron'') for wing. The Ancient Greek ὑμήν (''hymen'') for membrane provides a plausible etymology for the term because species in this order have membranous wings. However, a key characteristic of this order is that the hindwings are conn ...
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Triassic
The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. The Triassic Period is subdivided into three epochs: Early Triassic, Middle Triassic and Late Triassic. The Triassic began in the wake of the Permian–Triassic extinction event, which left the Earth's biosphere impoverished; it was well into the middle of the Triassic before life recovered its former diversity. Three categories of organisms can be distinguished in the Triassic record: survivors from the extinction event, new groups that flourished briefly, and other new groups that went on to dominate the Mesozoic Era. Reptiles, especially archosaurs, were the chief terrestrial vertebrates during this time. A specialized subgroup of archo ...
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Holometabolism
Holometabolism, also called complete metamorphosis, is a form of insect development which includes four life stages: egg, larva, pupa, and imago (or adult). Holometabolism is a synapomorphic trait of all insects in the superorder Endopterygota. Immature stages of holometabolous insects are very different from the mature stage. In some species the holometabolous life cycle prevents larvae from competing with adults because they inhabit different ecological niches. The morphology and behavior of each stage are adapted for different activities. For example, larval traits maximize feeding, growth, and development, while adult traits enable dispersal, mating, and egg laying. Some species of holometabolous insects protect and feed their offspring. Other insect developmental strategies include ametabolism and hemimetabolism. Developmental stages There are four general developmental stages, each with its own morphology and function. Egg The first stage of the insect life cycle is t ...
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Diptera
Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced mechanosensory organs known as halteres, which act as high-speed sensors of rotational movement and allow dipterans to perform advanced aerobatics. Diptera is a large order containing an estimated 1,000,000 species including horse-flies, crane flies, hoverflies and others, although only about 125,000 species have been described. Flies have a mobile head, with a pair of large compound eyes, and mouthparts designed for piercing and sucking (mosquitoes, black flies and robber flies), or for lapping and sucking in the other groups. Their wing arrangement gives them great maneuverability in flight, and claws and pads on their feet enable them to cling to smooth surfaces. Flies undergo complete metamorphosis; the eggs are often laid on the l ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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Protein
Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, responding to stimuli, providing structure to cells and organisms, and transporting molecules from one location to another. Proteins differ from one another primarily in their sequence of amino acids, which is dictated by the nucleotide sequence of their genes, and which usually results in protein folding into a specific 3D structure that determines its activity. A linear chain of amino acid residues is called a polypeptide. A protein contains at least one long polypeptide. Short polypeptides, containing less than 20–30 residues, are rarely considered to be proteins and are commonly called peptides. The individual amino acid residues are bonded together by peptide bonds and adjacent amino acid residues. The sequence of amino acid residue ...
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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about ance ...
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Melissa (mythology)
In Greek mythology, Melissa (Ancient Greek: Μέλισσα) may refer to the following women: * Melissa, a nymph who discovered and taught the use of honey, and from whom bees were believed to have received their name, μέλισσαι.Scholia ad Pindar, ''Pythian Odes'' 4.104 Bees seem to have been the symbol of nymphs, whence they themselves are sometimes called Melissae, and are sometimes said to have been metamorphosed into bees. Hence also nymphs in the form of bees are said to have guided the colonists that went to Ephesus; and the nymphs who nursed the infant Zeus are called Melissae, or Meliae. * Melissa, daughter of the Cretan king Melissus, who, together with her sister Amalthea, fed Zeus with goats' milk. She may be the same with the above Melissa. * Melissa, daughter of Epidamnus and mother of Dyrrhachius by Poseidon. Her father and son gave their name to the town in Illyria which was called Epidamnos and later on Dyrrhachium. The name Melissae was transferred to p ...
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Hymen (god)
Hymen ( grc, Ὑμήν), Hymenaios or Hymenaeus, in Hellenistic religion, is a god of marriage ceremonies, inspiring feasts and song. Related to the god's name, a ''hymenaios'' is a genre of Greek lyric poetry sung during the procession of the bride to the groom's house in which the god is addressed, in contrast to the ''Epithalamium'', which is sung at the nuptial threshold. He is one of the winged love gods, the Erotes. Hymen is the son of Apollo and one of the muses, Clio or Calliope or Urania or Terpsichore. Etymology Hymen's name is derived from the Proto-Indo-European root *''syuh₁-men''-, "to sew together," hence, "joiner;" it is also recorded in Doric Greek as Ῡ̔μᾱ́ν (''Hyman''). The term ''hymen'' was also used for a thin skin or membrane, such as the hymen that covers the vaginal opening and was traditionally supposed to be broken by sexual intercourse following a woman's (first) marriage. So, the membrane's name was not directly connected to that of the g ...
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Hamuli
A hamus or hamulus is a structure functioning as, or in the form of, hooks or hooklets. Etymology The terms are directly from Latin, in which ''hamus'' means "hook". The plural is ''hami''. ''Hamulus'' is the diminutive – hooklet or little hook. The plural is ''hamuli''. Adjectives are ''hamate'' and ''hamulate'', as in "a hamulate wing-coupling", in which the wings of certain insects in flight are joined by hooking hamuli on one wing into folds on a matching wing. ''Hamulate'' can also mean "having hamuli". The terms ''hamose'', ''hamular'', ''hamous'' and ''hamiform'' also have been used to mean "hooked", or "hook-shaped". Terms such as ''hamate'' that do not indicate a diminutive usually refer particularly to a hook at the tip, whereas diminutive terms such as ''hamulose'' tend to imply that something is beset with small hooks. Anatomy In vertebrate anatomy, a hamulus is a small, hook-shaped portion of a bone, or possibly of other hard tissue. In human anatomy, exampl ...
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Insect Wing
Insect wings are adult outgrowths of the insect exoskeleton that enable insects to fly. They are found on the second and third thoracic segments (the mesothorax and metathorax), and the two pairs are often referred to as the forewings and hindwings, respectively, though a few insects lack hindwings, even rudiments. The wings are strengthened by a number of longitudinal veins, which often have cross-connections that form closed "cells" in the membrane (extreme examples include the dragonflies and lacewings). The patterns resulting from the fusion and cross-connection of the wing veins are often diagnostic for different evolutionary lineages and can be used for identification to the family or even genus level in many orders of insects. Physically, some insects move their flight muscles directly, others indirectly. In insects with direct flight, the wing muscles directly attach to the wing base, so that a small downward movement of the wing base lifts the wing itself upward. Those i ...
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Etymology
Etymology ()The New Oxford Dictionary of English (1998) – p. 633 "Etymology /ˌɛtɪˈmɒlədʒi/ the study of the class in words and the way their meanings have changed throughout time". is the study of the history of the Phonological change, form of words and, by extension, the origin and evolution of their semantic meaning across time. It is a subfield of historical linguistics, and draws upon comparative semantics, Morphology_(linguistics), morphology, semiotics, and phonetics. For languages with a long recorded history, written history, etymologists make use of texts, and texts about the language, to gather knowledge about how words were used during earlier periods, how they developed in Semantics, meaning and Phonological change, form, or when and how they Loanword, entered the language. Etymologists also apply the methods of comparative linguistics to reconstruct information about forms that are too old for any direct information to be available. By analyzing related ...
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