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Horseshoe Crab
Horseshoe crabs are marine and brackish water arthropods of the family Limulidae and the only living members of the order Xiphosura. Despite their name, they are not true crabs or crustaceans: they are chelicerates, most closely related to arachnids, such as spiders and scorpions. Horseshoe crabs live primarily in and around shallow coastal waters on soft, sandy or muddy bottoms. They are generally found in the intertidal zone at spring high tides. They are eaten in some parts of Asia, and used as fishing bait, in fertilizer and in science (especially ''Limulus'' amebocyte lysate). In recent years, population declines have occurred as a consequence of coastal habitat destruction and overharvesting. Tetrodotoxin may be present in one horseshoe crab species, '' Carcinoscorpius rotundicauda''. Fossil records for horseshoe crabs extend back as far as 480 million years ago, with extant forms being living fossils. A 2019 molecular analysis places them as the sister group of Ricinule ...
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William Elford Leach
William Elford Leach Royal Society, FRS (2 February 1791 – 25 August 1836) was an English zoologist and marine biologist. Life and work Elford Leach was born at Hoe Gate, Plymouth, the son of an attorney. At the age of twelve he began a medical apprenticeship at the Royal Devon and Exeter Hospital, Devonshire and Exeter Hospital, studying anatomy and chemistry. By this time he was already collecting marine animals from Plymouth Sound and along the Devon coast. At seventeen he began studying medicine at St Bartholomew's Hospital in London, finishing his training at the University of Edinburgh before graduating Doctor of Medicine, MD from the University of St Andrews (where he had never studied). From 1813 Leach concentrated on his zoological interests and was employed as an 'Assistant Librarian' (what would later be called Assistant Keeper) in the Natural History Museum, London, Natural History Department of the British Museum, where he had responsibility for the zoological ...
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Carcinoscorpius Rotundicauda
The mangrove horseshoe crab (''Carcinoscorpius rotundicauda''), also known as the round-tailed horseshoe crab, is a chelicerate arthropod found in tropical marine and brackish waters in India, Bangladesh, Indonesia, Malaysia, Brunei, Singapore, Thailand, Cambodia, Vietnam, China and Hong Kong. It may also occur in Sri Lanka, Myanmar and the Philippines, but confirmed records are lacking. Despite their name, horseshoe crabs are more closely related to spiders and scorpions (all are in the subphylum Chelicerata) than to crabs. Recent phylogenetic analysis suggests that horseshoe crabs may themselves be arachnids. The mangrove horseshoe crab is the only species in the genus ''Carcinoscorpius''. There are four extant (living) species of horseshoe crab. The biology, ecology and breeding patterns of ''C. rotundicauda'' and the two other Asian horseshoe crab species, ''Tachypleus gigas'' and ''Tachypleus tridentatus'', have not been as well documented as those of the North American speci ...
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Species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, morphology, behaviour or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. However, only about 14% of these had been described by 2011. All species (except viruses) are given a two-part name, a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature, also sometimes i ...
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Order (biology)
Order ( la, wikt:ordo#Latin, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between Family_(biology), family and Class_(biology), class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. An immediately higher rank, superorder, is sometimes added directly above order, with suborder directly beneath order. An order can also be defined as a group of related families. What does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with a capital letter. Fo ...
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Family (biology)
Family ( la, familia, plural ') is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". What belongs to a family—or if a described family should be recognized at all—are proposed and determined by practicing taxonomists. There are no hard rules for describing or recognizing a family, but in plants, they can be characterized on the basis of both vegetative and reproductive features of plant species. Taxonomists often take different positions about descriptions, and there may be no broad consensus across the scientific community for some time. The publishing of new data and opini ...
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Ordovician
The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. The Ordovician, named after the Celtic Britons, Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same Rock (geology), rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed Stratum, strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Union of Geological Sciences, Intern ...
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Stratum
In geology and related fields, a stratum ( : strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as either '' bedding surfaces'' or ''bedding planes''.Salvador, A. ed., 1994. ''International stratigraphic guide: a guide to stratigraphic classification, terminology, and procedure. 2nd ed.'' Boulder, Colorado, The Geological Society of America, Inc., 215 pp. . Prior to the publication of the International Stratigraphic Guide, older publications have defined a stratum as either being either equivalent to a single bed or composed of a number of beds; as a layer greater than 1 cm in thickness and constituting a part of a bed; or a general term that includes both ''bed'' and ''lamina''.Neuendorf, K.K.E., Mehl, Jr., J.P., and Jackson, J.A. , eds., 2005. ''Glossary of Geology'' 5th ed. Alexandria, Virginia, American Geological Institute. 779 pp. . ...
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Fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the ...
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Chasmataspidid
Chasmataspidids, sometime referred to as chasmataspids, are a group of extinct chelicerate arthropods that form the order Chasmataspidida. Chasmataspidids are probably related to horseshoe crabs (Xiphosura) and/or sea scorpions (Eurypterida), with more recent studies suggest that they form a clade (Dekatriata) with Eurypterida and Arachnida. Chasmataspidids are known sporadically in the fossil record through to the mid-Devonian, with possible evidence suggesting that they were also present during the late Cambrian. Chasmataspidids are most easily recognised by having an opisthosoma divided into a wide forepart (preabdomen) and a narrow hindpart (postabdomen) each comprising 4 and 9 segments respectively. There is some debate about whether they form a natural (i.e. monophyletic) group. Distribution Chasmataspidids survived at least since Ordovician to mid-Devonian in age. As of 2019, most chasmataspidids (with a total of 9 species) are known from the Devonian strata, while the p ...
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Zeitschrift Für Zoologische Systematik Und Evolutionsforschung
''Journal of Zoological Systematics and Evolutionary Research'' (') is a quarterly, peer reviewed, scientific journal, published by Wiley-Blackwell. It was originally established in 1963, then reestablished in 1994 by John Wiley & Sons (some division of Blackwell). The editor in chief is Dr. Wilfried Westheide (, Germany). According to ''Journal Citation Reports'', the 2016 impact factor for this journal is 2.444. Scope The focus of ''Journal of Zoological Systematics and Evolutionary Research'' is the systematic study of animal sciences, connected with evolutionary research. Its function is both as a forum and a survey for the range of evolutionary research studies and related fields, which is published in English, German or French. A component of the focus is integrating original research results from anatomy, morphology, physiology, ethology, general genetics, population genetics, developmental biology, and molecular biology. Besides original research articles (20 pages) ...
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Eurypterid
Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the Order (biology), order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 Myr, million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic Chelicerata, chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus ''Eurypterus'' accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Tri ...
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Crustacean
Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group can be treated as a subphylum under the clade Mandibulata. It is now well accepted that the hexapods emerged deep in the Crustacean group, with the completed group referred to as Pancrustacea. Some crustaceans (Remipedia, Cephalocarida, Branchiopoda) are more closely related to insects and the other hexapods than they are to certain other crustaceans. The 67,000 described species range in size from '' Stygotantulus stocki'' at , to the Japanese spider crab with a leg span of up to and a mass of . Like other arthropods, crustaceans have an exoskeleton, which they moult to grow. They are distinguished from other groups of arthropods, such as insects, myriapods and chelicerates, by the possession of biramous (two-parted) limbs, and by th ...
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