Herkogamy
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Herkogamy
Herkogamy (or hercogamy) is the spatial separation of the anthers and stigma in hermaphroditic angiosperms. It is a common strategy for reducing self-fertilization. Common forms *Approach herkogamy - (called "pin flowers") is displayed when the stigma is displayed above the level of the anthers. This arrangement of sex organs causes floral visitors to first contact the stigma, before removing pollen from the anthers. This form of herkogamy is considered to be common, and is associated with a large, diverse fauna of floral visitors/pollinators. *Reverse herkogamy - (called "thrum flowers") is displayed when the stigma is recessed below the level of the anthers. This arrangement causes floral visitors to first contact the anthers before the stigma. For this reason, reverse herkogamy is believed to facilitate greater pollen export than approach herkogamy. This type of sex-organ arrangement is typically associated with Lepidopteran (moth or butterfly) pollination. See also * Heteros ...
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Heterostyly
Heterostyly is a unique form of polymorphism and herkogamy in flowers. In a heterostylous species, two or three morphological types of flowers, termed "morphs", exist in the population. On each individual plant, all flowers share the same morph. The flower morphs differ in the lengths of the pistil and stamens, and these traits are not continuous. The morph phenotype is genetically linked to genes responsible for a unique system of self-incompatibility, termed heteromorphic self-incompatibility, that is, the pollen from a flower on one morph cannot fertilize another flower of the same morph. Heterostylous plants having two flower morphs are termed " distylous". In one morph (termed "pin", "longistylous", or "long-styled" flower) the stamens are short and the pistils are long; in the second morph (termed "thrum", "brevistylous", or "short-styled" flower) the stamens are long and the pistils are short; the length of the pistil in one morph equals the length of the stamens in the ...
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Anther
The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament and an anther which contains ''microsporangia''. Most commonly anthers are two-lobed and are attached to the filament either at the base or in the middle area of the anther. The sterile tissue between the lobes is called the connective, an extension of the filament containing conducting strands. It can be seen as an extension on the dorsal side of the anther. A pollen grain develops from a microspore in the microsporangium and contains the male gametophyte. The stamens in a flower are collectively called the androecium. The androecium can consist of as few as one-half stamen (i.e. a single locule) as in '' Canna'' species or as many as 3,482 stamens which have been counted in the saguaro (''Carnegiea gigantea''). The androecium in var ...
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Stigma (botany)
The stigma () is the receptive tip of a carpel, or of several fused carpels, in the gynoecium of a flower. Description The stigma, together with the style and ovary (typically called the stigma-style-ovary system) comprises the pistil, which is part of the gynoecium or female reproductive organ of a plant. The stigma itself forms the distal portion of the style, or stylodia, and is composed of , the cells of which are receptive to pollen. These may be restricted to the apex of the style or, especially in wind pollinated species, cover a wide surface. The stigma receives pollen and it is on the stigma that the pollen grain germinates. Often sticky, the stigma is adapted in various ways to catch and trap pollen with various hairs, flaps, or sculpturings. The pollen may be captured from the air (wind-borne pollen, anemophily), from visiting insects or other animals ( biotic pollination), or in rare cases from surrounding water (hydrophily). Stigma can vary from long and sle ...
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Angiosperms
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants that produce their seeds enclosed within a fruit. They are by far the most diverse group of land plants with 64 orders, 416 families, approximately 13,000 known genera and 300,000 known species. Angiosperms were formerly called Magnoliophyta (). Like gymnosperms, angiosperms are seed-producing plants. They are distinguished from gymnosperms by characteristics including flowers, endosperm within their seeds, and the production of fruits that contain the seeds. The ancestors of flowering plants diverged from the common ancestor of all living gymnosperms before the end of the Carboniferous, over 300 million years ago. The closest fossil relatives of flowering plants are uncertain and contentious. The earliest angiosperm fossils are in the ...
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Pollen
Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophytes during the process of their movement from the stamens to the pistil of flowering plants, or from the male cone to the female cone of gymnosperms. If pollen lands on a compatible pistil or female cone, it germinates, producing a pollen tube that transfers the sperm to the ovule containing the female gametophyte. Individual pollen grains are small enough to require magnification to see detail. The study of pollen is called palynology and is highly useful in paleoecology, paleontology, archaeology, and forensics. Pollen in plants is used for transferring haploid male genetic material from the anther of a single flower to the stigma of another in cross-pollination. In a case of self-pollination, this process takes place from the anth ...
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Pollinator
A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are the major pollinators of most plants, and insect pollinators include all families of bees and most families of aculeate wasps; ants; many families of flies; many lepidopterans (both butterflies and moths); and many families of beetles. Vertebrates, mainly bats and birds, but also some non-bat mammals (monkeys, lemurs, possums, rodents) and some lizards pollinate certain plants. Among the pollinating birds are hummingbirds, honeyeaters and sunbirds with long beaks; they pollinate a number of deep-throated flowers. Humans may also carry out artificial pollination. A pollinator is different from a pollenizer, a plant that is a source of pollen for the pollination process. Background Plants fall into pollination syndromes that reflect the type o ...
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Lepidopteran
Lepidoptera ( ) is an order of insects that includes butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 families and 46 superfamilies, 10 percent of the total described species of living organisms. It is one of the most widespread and widely recognizable insect orders in the world. The Lepidoptera show many variations of the basic body structure that have evolved to gain advantages in lifestyle and distribution. Recent estimates suggest the order may have more species than earlier thought, and is among the four most speciose orders, along with the Hymenoptera, Diptera, and Coleoptera. Lepidopteran species are characterized by more than three derived features. The most apparent is the presence of scales that cover the bodies, wings, and a proboscis. The scales are modified, flattened "hairs", and give butterflies and moths their wide variety of colors and patterns. Almost all species have some form of membran ...
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Pollination
Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, and bats; water; wind; and even plants themselves, when self-pollination occurs within a closed flower. Pollination often occurs within a species. When pollination occurs between species, it can produce hybrid offspring in nature and in plant breeding work. In angiosperms, after the pollen grain (gametophyte) has landed on the stigma, it germinates and develops a pollen tube which grows down the style until it reaches an ovary. Its two gametes travel down the tube to where the gametophyte(s) containing the female gametes are held within the carpel. After entering an ovum cell through the micropyle, one male nucleus fuses with the polar bodies to produce the endosperm tissues, while the other fuses with the ovule to produce the embr ...
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