Heliophora Orbiculus
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Heliophora Orbiculus
''Heliophora orbicularis'', also known as the West African Sand Dollar, is a small sand dollar in to the family Rotulidae, and the only species in the genus ''Heliophora''. It, and other members of Rotulidae have been found in West African marine strata from the Late Miocene onward. Like the related , it is still extant. Description Individuals of ''H. orbicularis'' are very distinctive in appearance and also highly morphic, with a remarkable diversity in individual specimens. This genus has an extremely variable circular to oval-shaped test. The indentations may remain restricted to the posterior edge or may be found along the entire margin of the test. The indentations may be very shallow or quite deep, forming long "fingers," or digits. The digits are very fragile, and prone to breaking off. Ecology and feeding ''Heliophora orbicularis'' live in large groups in very shallow water of lagoons and estuaries, usually lightly buried in mud. Food particles are picked out of t ...
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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Senegal
Senegal,; Wolof: ''Senegaal''; Pulaar: 𞤅𞤫𞤲𞤫𞤺𞤢𞥄𞤤𞤭 (Senegaali); Arabic: السنغال ''As-Sinighal'') officially the Republic of Senegal,; Wolof: ''Réewum Senegaal''; Pulaar : 𞤈𞤫𞤲𞤣𞤢𞥄𞤲𞤣𞤭 𞤅𞤫𞤲𞤫𞤺𞤢𞥄𞤤𞤭 (Renndaandi Senegaali); Arabic: جمهورية السنغال ''Jumhuriat As-Sinighal'') is a country in West Africa, on the Atlantic Ocean coastline. Senegal is bordered by Mauritania to the north, Mali to the east, Guinea to the southeast and Guinea-Bissau to the southwest. Senegal nearly surrounds the Gambia, a country occupying a narrow sliver of land along the banks of the Gambia River, which separates Senegal's southern region of Casamance from the rest of the country. Senegal also shares a maritime border with Cape Verde. Senegal's economic and political capital is Dakar. Senegal is notably the westernmost country in the mainland of the Old World, or Afro-Eurasia. It owes its name to the ...
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Ambulacral
Ambulacral is a term typically used in the context of anatomical parts of the phylum Echinodermata or class Sea star, Asteroidea and Edrioasteroidea. Echinoderms can have ambulacral parts that include Ossicle (echinoderm), ossicles, plates, spines, and suckers. For example, sea stars or "star fish" have an ambulacral groove on their oral side (underside). This ambulacral groove extends from the mouth to the end of each ray or arm. Each groove of each arm in turn has four rows of hollow tube feet that can be extended or withdrawn. Opposite the ambulacral groove is an ambulacral ridge on the aboral side of each ray, known as an ambulacrum (anatomy), ambulacrum. These have interambulacra between them. Etymology From the Latin 'ambulācrum', meaning 'walk planted with trees', 'avenue', 'alley' and 'walking place' Derives from' 'ambulāre', meaning 'to walk' or 'Amble' meaning 'To walk slowly or leisurely'. Has Indo-European roots - deriving from 'Ambhi'The Free Dictionary (n.d.) amb ...
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Blastopore
Gastrulation is the stage in the early embryonic development of most animals, during which the blastula (a single-layered hollow sphere of cells), or in mammals the blastocyst is reorganized into a multilayered structure known as the gastrula. Before gastrulation, the embryo is a continuous epithelial sheet of cells; by the end of gastrulation, the embryo has begun differentiation to establish distinct cell lineages, set up the basic axes of the body (e.g. dorsal-ventral, anterior-posterior), and internalized one or more cell types including the prospective gut. In triploblastic organisms, the gastrula is trilaminar (three-layered). These three germ layers are the ectoderm (outer layer), mesoderm (middle layer), and endoderm (inner layer).Mundlos 2009p. 422/ref>McGeady, 2004: p. 34 In diploblastic organisms, such as Cnidaria and Ctenophora, the gastrula has only ectoderm and endoderm. The two layers are also sometimes referred to as the ''hypoblast'' and ''epiblast''. Sponges ...
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Deuterostome
Deuterostomia (; in Greek) are animals typically characterized by their anus forming before their mouth during embryonic development. The group's sister clade is Protostomia, animals whose digestive tract development is more varied. Some examples of deuterostomes include vertebrates (and thus humans), sea stars, and crinoids. In deuterostomy, the developing embryo's first opening (the blastopore) becomes the anus, while the mouth is formed at a different site later on. This was initially the group's distinguishing characteristic, but deuterostomy has since been discovered among protostomes as well. This group is also known as enterocoelomates, because their coelom develops through enterocoely. The three major clades of deuterostomes are Chordata (e.g. vertebrates), Echinodermata (e.g. starfish), and Hemichordata (e.g. acorn worms). Together with Protostomia and their out-group Xenacoelomorpha, these compose the Bilateria, animals with bilateral symmetry and three germ layers. ...
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Ciliated
The cilium, plural cilia (), is a membrane-bound organelle found on most types of eukaryotic cell, and certain microorganisms known as ciliates. Cilia are absent in bacteria and archaea. The cilium has the shape of a slender threadlike projection that extends from the surface of the much larger cell body. Eukaryotic flagella found on sperm cells and many protozoans have a similar structure to motile cilia that enables swimming through liquids; they are longer than cilia and have a different undulating motion. There are two major classes of cilia: ''motile'' and ''non-motile'' cilia, each with a subtype, giving four types in all. A cell will typically have one primary cilium or many motile cilia. The structure of the cilium core called the axoneme determines the cilium class. Most motile cilia have a central pair of single microtubules surrounded by nine pairs of double microtubules called a 9+2 axoneme. Most non-motile cilia have a 9+0 axoneme that lacks the central pair of mi ...
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Endoskeleton
An endoskeleton (From Greek ἔνδον, éndon = "within", "inner" + σκελετός, skeletos = "skeleton") is an internal support structure of an animal, composed of mineralized tissue. Overview An endoskeleton is a skeleton that is on the inside of a body, like humans, dogs, or some fish. The endoskeleton develops within the skin or in the deeper body tissues. The vertebrate endoskeleton is basically made up of two types of tissues (bone and cartilage). During early embryonic development the endoskeleton is composed of notochord and cartilage. The notochord in most vertebrates is replaced by the vertebral column and cartilage is replaced by bone in most adults. In three phyla and one subclass of animals, endoskeletons of various complexity are found: Chordata, Echinodermata, Porifera, and Coleoidea. An endoskeleton may function purely for support (as in the case of sponges), but often serves as an attachment site for muscle and a mechanism for transmitting muscular forces ...
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Ossicle (echinoderm)
Ossicles are small calcareous elements embedded in the dermis of the body wall of echinoderms. They form part of the endoskeleton and provide rigidity and protection. They are found in different forms and arrangements in sea urchins, starfish, brittle stars, sea cucumbers, and crinoids. The ossicles and spines (which are specialised sharp ossicles) are the only parts of the animal likely to be fossilized after an echinoderm dies. Formation Ossicles are created intracellularly by specialised secretory cells known as sclerocytes in the dermis of the body wall of echinoderms. Each ossicle is composed of microcrystals of calcite arranged in a three-dimensional lattice known as a stereom. Under polarized light the ossicle behaves as if it were a single crystal because the axes of all the crystals are parallel. The space between the crystals is known as the stroma and allows entry to sclerocytes for enlargement and repair. The honeycomb structure is light but tough and collage ...
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Calcareous
Calcareous () is an adjective meaning "mostly or partly composed of calcium carbonate", in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines. In zoology ''Calcareous'' is used as an adjectival term applied to anatomical structures which are made primarily of calcium carbonate, in animals such as gastropods, i.e., snails, specifically about such structures as the operculum, the clausilium, and the love dart. The term also applies to the calcium carbonate tests of often more or less microscopic Foraminifera. Not all tests are calcareous; diatoms and radiolaria have siliceous tests. The molluscs are calcareous, as are calcareous sponges ( Porifera), that have spicules which are made of calcium carbonate. In botany ''Calcareous grassland'' is a form of grassland characteristic of soils containing much calcium carbonate from underlying chalk or limestone rock. In medicine The term is used in pathology, for example i ...
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Mesoderm
The mesoderm is the middle layer of the three germ layers that develops during gastrulation in the very early development of the embryo of most animals. The outer layer is the ectoderm, and the inner layer is the endoderm.Langman's Medical Embryology, 11th edition. 2010. The mesoderm forms mesenchyme, mesothelium, non-epithelial blood cells and coelomocytes. Mesothelium lines coeloms. Mesoderm forms the muscles in a process known as myogenesis, septa (cross-wise partitions) and mesenteries (length-wise partitions); and forms part of the gonads (the rest being the gametes). Myogenesis is specifically a function of mesenchyme. The mesoderm differentiates from the rest of the embryo through intercellular signaling, after which the mesoderm is polarized by an organizing center. The position of the organizing center is in turn determined by the regions in which beta-catenin is protected from degradation by GSK-3. Beta-catenin acts as a co-factor that alters the activity of ...
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Ectoderm
The ectoderm is one of the three primary germ layers formed in early embryonic development. It is the outermost layer, and is superficial to the mesoderm (the middle layer) and endoderm (the innermost layer). It emerges and originates from the outer layer of germ cells. The word ectoderm comes from the Greek ''ektos'' meaning "outside", and ''derma'' meaning "skin".Gilbert, Scott F. Developmental Biology. 9th ed. Sunderland, MA: Sinauer Associates, 2010: 333-370. Print. Generally speaking, the ectoderm differentiates to form epithelial and neural tissues (spinal cord, peripheral nerves and brain). This includes the skin, linings of the mouth, anus, nostrils, sweat glands, hair and nails, and tooth enamel. Other types of epithelium are derived from the endoderm. In vertebrate embryos, the ectoderm can be divided into two parts: the dorsal surface ectoderm also known as the external ectoderm, and the neural plate, which invaginates to form the neural tube and neural crest. Th ...
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Endoderm
Endoderm is the innermost of the three primary germ layers in the very early embryo. The other two layers are the ectoderm (outside layer) and mesoderm (middle layer). Cells migrating inward along the archenteron form the inner layer of the gastrula, which develops into the endoderm. The endoderm consists at first of flattened cells, which subsequently become columnar. It forms the epithelial lining of multiple systems. In plant biology, endoderm corresponds to the innermost part of the cortex ( bark) in young shoots and young roots often consisting of a single cell layer. As the plant becomes older, more endoderm will lignify. Production The following chart shows the tissues produced by the endoderm. The embryonic endoderm develops into the interior linings of two tubes in the body, the digestive and respiratory tube. Liver and pancreas cells are believed to derive from a common precursor. In humans, the endoderm can differentiate into distinguishable organs after 5 week ...
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