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Haplochromine
__NOTOC__ The haplochromine cichlids are a tribe of cichlids in subfamily Pseudocrenilabrinae called Haplochromini. This group includes the type genus (''Haplochromis'') plus a number of closely related genera such as '' Aulonocara'', '' Astatotilapia'', and '' Chilotilapia''. They are endemic to eastern, southern and northern Africa, except for ''Astatotilapia flaviijosephi'' in the ''Middle East''. A common name in a scientific context is East African cichlids – while they are not restricted to that region, they are the dominant Cichlidae there. This tribe was extensively studied by Ethelwynn Trewavas, who made major reviews in 1935 and 1989, at the beginning and at the end of her career in ichthyology. Even today, numerous new species are being described each year. The haplochromines were in older times treated as subfamily Haplochrominae, However, the great African radiation of pseudocrenilabrine cichlids is certainly not monophyletic without them, and thus they are t ...
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Cichlid
Cichlids are fish from the family Cichlidae in the order Cichliformes. Cichlids were traditionally classed in a suborder, the Labroidei, along with the wrasses ( Labridae), in the order Perciformes, but molecular studies have contradicted this grouping. The closest living relative of cichlids is probably the convict blenny, and both families are classified in the 5th edition of ''Fishes of the World'' as the two families in the Cichliformes, part of the subseries Ovalentaria. This family is both large and diverse. At least 1,650 species have been scientifically described, making it one of the largest vertebrate families. New species are discovered annually, and many species remain undescribed. The actual number of species is therefore unknown, with estimates varying between 2,000 and 3,000. Many cichlids, particularly tilapia, are important food fishes, while others, such as the ''Cichla'' species, are valued game fish. The family also includes many popular freshwater aquariu ...
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Haplochromis
''Haplochromis'' is a ray-finned fish genus in the family Cichlidae. It has been used as the default "wastebin taxon" for Pseudocrenilabrinae cichlids of the East African Rift, and as such became the "largest" fish "genus". Many of these cichlids are popular aquarium fishes; like similar Haplochromini they are known as "haplos", "happies" or "haps" among aquarium enthusiasts. The genus was established by F.M. Hilgendorf in 1888. It was originally conceived as a subgenus of A.C.L.G. Günther's ''"Chromis"'', at that time an even larger "wastebin genus" for Pseudocrenilabrinae cichlids. The type species of Hilgendorf was '' H. obliquidens''. ''"Chromis"'' of Günther turned out to be a junior homonym of G. Cuvier's ocean fish genus '' Chromis'', already established in 1814, and was abolished. As the years went by, other genera of (mostly) Haplochromini were lumped with and split again from ''Haplochromis'', and the final delimitation of the clade around ''H. obliquidens'' is not ...
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Astatotilapia Flaviijosephi
''Astatotilapia flaviijosephi'', the Jordan mouthbrooder, is a vulnerable species of freshwater fish in the family Cichlidae (cichlids). It is found in the central Jordan River system, including Lake Tiberias (Kinneret), in Israel, Jordan and Syria, making it the only haplochromine cichlid to naturally range outside of Africa. This species is too small to be of significant importance to fisheries, unlike the only other cichlids native to the Levant, the economically important tilapias (''Oreochromis aureus'', '' O. niloticus'', ''Sarotherodon galilaeus'', ''Coptodon zillii'' and ''Tristramella''). The specific name ''flaviijosephi'' refers to the historian Titus Flavius Josephus (37–c. 100 CE). Habitat and conservation status The natural habitats of ''A. flaviijosephi'' are streams, springs, canals, pools and shallow waters in lakes, especially in areas with stones or aquatic vegetation. The various subpopulations are isolated from each other, with some being lacustrine an ...
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Adaptive Radiation
In evolutionary biology, adaptive radiation is a process in which organisms diversify rapidly from an ancestral species into a multitude of new forms, particularly when a change in the environment makes new resources available, alters biotic interactions or opens new environmental niches. Starting with a single ancestor, this process results in the speciation and phenotypic adaptation of an array of species exhibiting different morphological and physiological traits. The prototypical example of adaptive radiation is finch speciation on the Galapagos ("Darwin's finches"), but examples are known from around the world. Characteristics Four features can be used to identify an adaptive radiation: #A common ancestry of component species: specifically a ''recent'' ancestry. Note that this is not the same as a monophyly in which ''all'' descendants of a common ancestor are included. #A phenotype-environment correlation: a ''significant'' association between environments and the morp ...
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Aulonocara
''Aulonocara'' is a genus of haplochromine cichlids endemic to Lake Malawi in East Africa. All ''Aulonocara'' species are maternal mouth brooders. Particularly in the aquarium hobby, ''Aulonocara'' species are also known as peacock cichlids, aulonocaras or simply "peacocks". This genus is strongly sexually dichromic, even by haplochromine standards. As aquarium fish, they are best kept with other medium-sized nonaggressive cichlids from Lake Malawi. All peacock cichlids are known to be less aggressive than their Mbuna counterparts, and thrive in warmer waters that have a slightly basic pH. Species The 22 recognized species in this genus are: * '' Aulonocara aquilonium'' Konings, 1995 * '' Aulonocara auditor'' ( Trewavas, 1935) * '' Aulonocara baenschi'' M. K. Meyer & Riehl, 1985 (nkhomo-benga peacock) * '' Aulonocara brevinidus'' Konings, 1995 * '' Aulonocara brevirostre'' ( Trewavas, 1935) * '' Aulonocara ethelwynnae'' M. K. Meyer, Riehl & Zetzsche, 1987 (Chitande aul ...
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Astatotilapia
''Astatotilapia'' is a genus of small freshwater fish in the family Cichlidae found in Eastern and Northern Africa, with a single species, ''A. flaviijosephi'', in Western Asia (the only non-African haplochromine). Many species have been moved between this genus and ''Haplochromis'', and while some consensus has been reached in recent years, their mutual delimitation is still far from settled. Based on mtDNA, ''Astatotilapia'' as currently defined is polyphyletic.Genner; Ngatunga; Mzighani; Smith; and Turner (2015). Geographical ancestry of Lake Malawi’s cichlid fish diversity. Biol. Lett. 11: 2015023. Species There are currently 10 recognized species in this genus: * ''Astatotilapia bloyeti'' ( Sauvage, 1883) (Bloyet's haplo) * ''Astatotilapia burtoni'' ( Günther, 1894) * ''Astatotilapia calliptera'' ( Günther, 1894) (Eastern happy) * ''Astatotilapia desfontainii'' ( Lacépède, 1802) * ''Astatotilapia flaviijosephi'' ( Lortet, 1883) (Jordan mouthbrooder) * ''Astatotilapia ...
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Copadichromis Cf
''Copadichromis'' is a genus of haplochromine cichlids endemic to Lake Malawi in Eastern Africa. ''Copadichromis'' are part of a group known as utaka and are popular with aquarists, as this genus is relatively peaceful in captivity compared to the mbuna. Species There are currently 25 recognized species in this genus: * '' Copadichromis atripinnis'' Stauffer & Te. Sato, 2002 * ''Copadichromis azureus'' Konings, 1990 * ''Copadichromis borleyi'' ( Iles, 1960) (redfin hap, goldfin hap) * ''Copadichromis chizumuluensis'' Stauffer & Konings, 2006 * ''Copadichromis chrysonotus'' (Boulenger, 1908) * '' Copadichromis cyaneus'' ( Trewavas, 1935) * ''Copadichromis cyanocephalus'' Stauffer & Konings, 2006 * ''Copadichromis diplostigma'' Stauffer & Konings, 2006 * ''Copadichromis geertsi'' Konings, 1999 * '' Copadichromis ilesi'' Konings, 1999 * ''Copadichromis insularis'' Stauffer & Konings, 2006 * ''Copadichromis jacksoni'' ( Iles, 1960) * ''Copadichromis likomae'' ( Iles, 1960) * ''C ...
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Mouthbrooding
Mouthbrooding, also known as oral incubation and buccal incubation, is the care given by some groups of animals to their offspring by holding them in the mouth of the parent for extended periods of time. Although mouthbrooding is performed by a variety of different animals, such as the Darwin's frog, fish are by far the most diverse mouthbrooders. Mouthbrooding has evolved independently in several different families of fish. Mouthbrooding behaviour Paternal mouthbrooders are species where the male looks after the eggs. Paternal mouthbrooders include the arowana, various mouthbrooding bettas and gouramies such as ''Betta pugnax'', and sea catfish such as ''Ariopsis felis''. Among cichlids, paternal mouthbrooding is relatively rare, but is found among some of the tilapiines, most notably the black-chin tilapia ''Sarotherodon melanotheron''. In the case of the maternal mouthbrooders, the female takes the eggs. Maternal mouthbrooders are found among both African and South American ci ...
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Astatotilapia Burtoni
''Astatotilapia burtoni'' is a species of fish in the family Cichlidae. It is found in Lake Tanganyika and its surrounding waterways, including parts of Burundi, Rwanda, Tanzania, and Zambia. Its natural habitats are rivers, intermittent rivers, swamps, freshwater lakes, freshwater marshes, intermittent freshwater marshes, and inland deltas. ''Astatotilapia burtoni'' has been used as a model organism to study the behaviors and physical systems of cichlids, including their development and embryogenesis. Moreover, the phylogenetic position of this particular species makes it an ideal model system for comparative genomic research. ''A. burtoni'' belongs under the haplochromines, which is the lineage of cichlids with the most species, and has been discovered to be a sister group to both the Lake Victoria region superflock (which has about 600 species) and the species flock of Lake Malawi (which has about 1,000 species). Social behavior Reversible male social roles The males of t ...
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Nimbochromis
''Nimbochromis'' is a small genus of haplochromine cichlids mostly endemic to Lake Malawi in East Africa. They are known as sleeper cichlids or ''kaligono'' ("sleepers" in Chichewa) due to their unique hunting behaviour. These piscivorous species are often seen lying motionless on the lake bottom near rocks where mbuna live, even adopting an unusual sideways position rarely seen in living fish. If smaller fishes approach, the ''Nimbochromis'' will "wake up" and try to seize them. Their coloration has an irregular dark cloudy pattern on lighter background; for one thing, this provides camouflage, but it is also suspected that it is – at least in some – evolving into aggressive mimicry (apparent death) by imitating a rotting fish carcass and thus luring scavengers to their demise. Species There are currently five recognized species in this genus: * '' Nimbochromis fuscotaeniatus'' (Regan, 1922) (Spothead Hap, Fuscotaeniatus Hap) * '' Nimbochromis linni'' ( W. E. Burges ...
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African Great Lakes
The African Great Lakes ( sw, Maziwa Makuu; rw, Ibiyaga bigari) are a series of lakes constituting the part of the Rift Valley lakes in and around the East African Rift. They include Lake Victoria, the second-largest fresh water lake in the world by area, Lake Tanganyika, the world's second-largest freshwater lake by volume and depth, and Lake Malawi, the world's eighth-largest fresh water lake by area. Collectively, they contain 31,000 km3 (7400 cu mi) of water, which is more than either Lake Baikal or the North American Great Lakes. This total constitutes about 25% of the planet's unfrozen surface fresh water. The large rift lakes of Africa are the ancient home of great biodiversity, and 10% of the world's fish species live in this region. Riparian countries in the African Great Lakes region include: Burundi, the Democratic Republic of the Congo, Ethiopia, Kenya, Malawi, Mozambique, Rwanda, Zambia, Tanzania, and Uganda. Lakes and drainage basins The following are ...
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Ethelwynn Trewavas
Ethelwynn Trewavas (5 November 1900 – 16 August 1993) was an ichthyologist at the British Museum of Natural History. She was known for her work on the families Cichlidae and Sciaenidae. She worked with Charles Tate Regan, another ichthyologist and taxonomist. Academic studies and career She received her bachelor's degree and Board of Education Certificate in Teaching in 1921 from Reading University, and then worked as a teacher before being employed by the King's College of Household and Social Science as a part-time demonstrator, spending most of her time on research. She was taught by Dr. Nellie B. Eales when associated with the Freshwater Biological Association. She met Charles Regan and was employed by him as his assistant until hired by the British Museum (Natural History) as Assistant Keeper in 1935. She was appointed Deputy Keeper of Zoology in 1958, and retired in 1961. She served as the senior scientist in the Fish Section of the British Museum (Natural Histo ...
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