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Graham Budd
Graham Edward Budd is a British palaeontologist. He is Professor and head of palaeobiology at Uppsala University. Budd's research focuses on the Cambrian explosion and on the evolution and development, anatomy, and patterns of diversification of the Ecdysozoa, a group of animals that include arthropods. Life and work Budd was born on 7 September 1968 in Colchester (Essex). He obtained his undergraduate degree at the University of Cambridge and remained there to continue his studies at a doctoral level by investigating the Sirius Passet fossil lagerstätte from the Cambrian of North Greenland. He finished his doctorate in 1994, with one of the findings being a new species of lobopodian, ''Kerygmachela''. Budd then moved to Sweden as a postdoc along with his PhD supervisor John Peel. Together with Sören Jensen he reintroduced the concepts of stem and crown groups to phylogenetics and is a major critic of molecular clocks current usage in determining the origin o ...
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Colchester
Colchester ( ) is a city in Essex, in the East of England. It had a population of 122,000 in 2011. The demonym is Colcestrian. Colchester occupies the site of Camulodunum, the first major city in Roman Britain and its first capital. Colchester therefore claims to be Britain's first city. It has been an important military base since the Roman era, with Colchester Garrison currently housing the 16th Air Assault Brigade. Situated on the River Colne, Colchester is northeast of London. The city is connected to London by the A12 road and the Great Eastern Main Line railway. Colchester is less than from London Stansted Airport and from the port of Harwich. Attractions in and around the city include Colchester United Football Club, Colchester Zoo, and several art galleries. Colchester Castle was constructed in the eleventh century on earlier Roman foundations; it now contains a museum. The main campus of the University of Essex is located just outside the city. Local governme ...
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Arthropod
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arthropod cuticle, cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an exoskeleton, external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior Organ (anatomy), organs; it has an open circulatory system. Like their exteriors, the internal or ...
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Acta Zoologica
''Acta Zoologica'' is a peer-reviewed scientific journal published by Wiley-Blackwell on behalf of the Royal Swedish Academy of Sciences and the Royal Danish Academy of Sciences and Letters. It was established in 1920 as ''Acta Zoologica'' and obtained its current name in 2002. It is one of the world's leading zoological journals and focuses on animal development, structure, and function, including physiological organization. It primarily publishes original research papers, but occasionally also publishes review articles. Increasingly, it has concentrated on animal development, with emphasis on the functional, comparative, and phylogenetic aspects. The editor-in-chief is Lennart Olsson (University of Jena). Abstracting and indexing The journal is abstracted and indexed in: According to the ''Journal Citation Reports'', the journal has a 2020 impact factor of 1.261, ranking it 15th out of 21 journals in the category "Anatomy & Morphology" and 105th out of 174 journals in the cat ...
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Molecular Clock
The molecular clock is a figurative term for a technique that uses the mutation rate of biomolecules to deduce the time in prehistory when two or more life forms diverged. The biomolecular data used for such calculations are usually nucleotide sequences for DNA, RNA, or amino acid sequences for proteins. The benchmarks for determining the mutation rate are often fossil or archaeological dates. The molecular clock was first tested in 1962 on the hemoglobin protein variants of various animals, and is commonly used in molecular evolution to estimate times of speciation or radiation. It is sometimes called a gene clock or an evolutionary clock. Early discovery and genetic equidistance The notion of the existence of a so-called "molecular clock" was first attributed to Émile Zuckerkandl and Linus Pauling who, in 1962, noticed that the number of amino acid differences in hemoglobin between different lineages changes roughly linearly with time, as estimated from fossil evidence. ...
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Phylogenetics
In biology, phylogenetics (; from Greek language, Greek wikt:φυλή, φυλή/wikt:φῦλον, φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, Protein, protein Amino acid, amino acid sequences, or Morphology (biology), morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An un ...
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Crown Group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crown gr ...
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Sören Jensen
Sören is a municipality in the district of Rendsburg-Eckernförde, in Schleswig-Holstein, Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe .... References Rendsburg-Eckernförde {{RendsburgEckernförde-geo-stub ...
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Postdoctoral Researcher
A postdoctoral fellow, postdoctoral researcher, or simply postdoc, is a person professionally conducting research after the completion of their doctoral studies (typically a PhD). The ultimate goal of a postdoctoral research position is to pursue additional research, training, or teaching in order to have better skills to pursue a career in academia, research, or any other field. Postdocs often, but not always, have a temporary academic appointment, sometimes in preparation for an academic faculty position. They continue their studies or carry out research and further increase expertise in a specialist subject, including integrating a team and acquiring novel skills and research methods. Postdoctoral research is often considered essential while advancing the scholarly mission of the host institution; it is expected to produce relevant publications in peer-reviewed academic journals or conferences. In some countries, postdoctoral research may lead to further formal qualificati ...
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Kerygmachela
''Kerygmachela kierkegaardi'' is a gilled lobopodian from the Cambrian Stage 3 aged Sirius Passet Lagerstätte in northern Greenland. Its anatomy strongly suggests that it, along with its relative ''Pambdelurion whittingtoni'', was a close relative of radiodont (anomalocaridids) and euarthropods. The generic name "Kerygmachela" derives from the Greek words ''Kerygma'' (proclamation) and ''Chela'' (claw), in reference to the flamboyant frontal appendages. The specific name, "kierkegaardi" honors Danish philosopher Søren Kierkegaard. Morphology The head of ''Kerygmachela'' possesses a pair of well-developed frontal appendages which correspond to those of other dinocaridids and siberiid lobopodians. Each of them terminates in a series of long spines. A pair of sessile, slit-like compound eyes is located slightly behind the base of these appendages. A small anterior-facing mouth is located below the head and bears a pair of stylet-like structures. The head also possesses a ...
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Lobopodia
The lobopodians, members of the informal group Lobopodia (from the Greek language, Greek, meaning "blunt feet"), or the formally erected phylum Lobopoda Cavalier-Smith (1998), are panarthropods with stubby legs called lobopods, a term which may also be used as a common name of this group as well. While the definition of lobopodians may differ between literatures, it usually refers to a group of soft-bodied, worm-like fossil panarthropods such as ''Aysheaia'' and ''Hallucigenia''. The oldest near-complete fossil lobopodians date to the Lower Cambrian; some are also known from Ordovician, Silurian and Carboniferous Lagerstätten. Some bear toughened claws, plates or spines, which are commonly preserved as small carbonaceous fossil, carbonaceous or Small Shelly Fossils, mineralized microfossils in Cambrian strata. The grouping is considered to be paraphyletic, as the three living panarthropod groups (Arthropoda, Tardigrada and Onychophora) are thought to have evolved from lobopodian ...
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North Greenland
The Northern Inspectorate of Greenland also known as North Greenland was a Danish inspectorate on Greenland consisting of the trading centers and missionary stations along the northwest coast of the island. History North Greenland was established in 1721. Its capital was at Godhavn (modern Qeqertarsuaq). The southernmost town of North Greenland was Egedesminde, which bordered Holsteinborg, which was the northernmost town of South Greenland. This boundary between North and South Greenland ran at around 68°N latitude, and in the North, North Greenland stretched to 78°N to enclose Thule. In 1911, as the administration of the colony was removed from the Royal Greenland Trading Department and folded into the Danish Ministry of the Interior, a provincial council ( da, landsråd) was established. It was elected indirectly from the local councils and had little say in the management of the colony. North Greenland was united with South Greenland in 1950,
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Cambrian
The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Ca ...
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