Gnatusuchus Pebasensis
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Gnatusuchus Pebasensis
''Gnatusuchus'' is an extinct genus of caiman represented by the type species ''Gnatusuchus pebasensis'' from the Middle Miocene Pebas Formation of Peru. ''Gnatusuchus'' lived about 13 million years ago (Ma) in a large wetland system called the Pebas mega-wetlands that covered over one million square kilometers of what is now the Amazon Basin (the modern basin had not yet developed at that time and instead of draining from west to east into the Atlantic Ocean, river systems drained northward through the wetlands and into the Caribbean Sea). Discovery and naming Fish and molluscs have long been known from the Pebas Formation, however starting in 2002 systematic surveys of the Peruvian Iquitos area have led to the discovery of many vertebrate remains including further fish remains, mammals, turtles and an abundance of crocodilians, with two contemporaneous lignitic bonebeds preserving a minimum of 7 coexisting taxa. Among the fossils recovered from the Iquitos localities is the holo ...
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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Simosuchus
''Simosuchus'' (meaning "pug-nosed crocodile" in Greek, referring to the animal's blunt snout) is an extinct genus of notosuchian crocodylomorphs from the Late Cretaceous of Madagascar. It is named for its unusually short skull. Fully grown individuals were about in length. The type species is ''Simosuchus clarki'', found from the Maevarano Formation in Mahajanga Province, although some fossils have been found in India. The teeth of ''S. clarki'' were shaped like maple leaves, which coupled with its short and deep snout suggests it was not a carnivore like most other crocodylomorphs. In fact, these features have led many palaeontologists to consider it a herbivore. Description ''Simosuchus'' was small, about long based on the skeletons of mature individuals. In contrast to most other crocodyliforms, which have long, low skulls, ''Simosuchus'' has a distinctively short snout. The snout resembles that of a pug, giving the genus its name, which means "pug-nosed crocodile" in Gre ...
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Brachychampsa
''Brachychampsa'' is an extinct genus of alligatoroid, possibly a basal caiman. Specimens have been reported from New Mexico, Colorado, Wyoming, Montana, North and South Dakota, New Jersey, and Saskatchewan, though only those from Montana, Utah, and New Mexico are based on material sufficient to justify the referral. One specimen has been reported from the Darbasa Formation of Kazakhstan, although the species status is indeterminate for the fossil. The genus first appeared during the late Campanian stage of the Late Cretaceous (Judithian North American stage) and became extinct during the late Maastrichtian stage of the Cretaceous (Lancian North American Land Mammal "Age"). ''Brachychampsa'' is distinguished by an enlarged fifth maxillary tooth in the upper jaw. Species The type species of ''Brachychampsa'' is ''B. montana'', first discovered from the Hell Creek Formation of Montana and described by Charles W. Gilmore in a paper in 1911. In that same paper, Gilmore recombine ...
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Albertochampsa
''Albertochampsa'' is an extinct genus of globidontan alligatoroid (possibly a stem-caiman) from the Late Cretaceous of Alberta. It was named in 1972 by Bruce Erickson, and the type species is ''A. langstoni''. It is known from a skull from the Campanian-age Dinosaur Park Formation, where it was rare; ''Leidyosuchus ''Leidyosuchus'' (meaning " Leidy's crocodile") is an extinct genus of alligatoroid from the Late Cretaceous of Alberta. It was named in 1907 by Lawrence Lambe, and the type species is ''L. canadensis''. It is known from a number of specimens fro ...'' is the most commonly found crocodilian at the Park. The skull of ''Albertochampsa'' was only about 21 cm long (8.3 in). References Crocodilians Late Cretaceous crocodylomorphs of North America Late Cretaceous reptiles of North America Fossil taxa described in 1972 Prehistoric pseudosuchian genera {{Cretaceous-reptile-stub ...
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Alligatorinae
Alligatorinae is a subfamily within the family Alligatoridae that contains the alligators and their closest extinct relatives, and is the sister taxon to Caimaninae (the caimans). Many genera in Alligatorinae are described, but only the genus ''Alligator'' is still living, with the remaining genera extinct. Evolution Alligators and caimans split in North America during the early Tertiary or late Cretaceous (about 53 million to about 65 million years ago). The Chinese alligator split from the American alligator about 33 million years ago and likely descended from a lineage that crossed the Bering land bridge during the Neogene. The modern American alligator is well represented in the fossil record of the Pleistocene. The alligator's full mitochondrial genome was sequenced in the 1990s. The full genome, published in 2014, suggests that the alligator evolved much more slowly than mammals and birds. Phylogeny Alligatorinae is cladistically defined as ''Alligator mississippiensi ...
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Globidonta
Globidonta is a clade of alligatoroids that includes alligators, caimans, and closely related extinct forms. It is defined as a stem-based clade including ''Alligator mississippiensis'' (the American Alligator) and all forms more closely related to it than to ''Diplocynodon''. The group's fossil range extends back into the Late Cretaceous with early alligatoroids such as '' Albertochampsa'' and '' Brachychampsa''. Extinct globidontans were particularly common in North America and Eurasia, and their modern range also includes South America. Basal globidontans are characterized by their blunt snouts and bulbous teeth. Modern globidontans have flattened snouts and more conical teeth, and are seen as more generalized than earlier globidontans. Generalized forms are usually expected to be ancestral to more specialized forms rather than descendants of them, so it is unusual for basal members of the group to appear specialized. This seems to conflict with the "Law of the Unspecialized ...
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Cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to descendants, nor does it show how much they have changed, so many differing evolutionary trees can be consistent with the same cladogram. A cladogram uses lines that branch off in different directions ending at a clade, a group of organisms with a last common ancestor. There are many shapes of cladograms but they all have lines that branch off from other lines. The lines can be traced back to where they branch off. These branching off points represent a hypothetical ancestor (not an actual entity) which can be inferred to exhibit the traits shared among the terminal taxa above it. This hypothetical ancestor might then provide clues about the order of evolution of various features, adaptation, and other evolutionary narratives about ance ...
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Globidentosuchus
''Globidentosuchus'' is an extinct genus of basal caimanine crocodylian known from the late Middle to Late Miocene of the Middle and the Upper Members of the Urumaco Formation at Urumaco, Venezuela. Its skull was very short and robust, with large units of spherical teeth used to break the shells of molluscs as part of its durophagus diet. It is thought to be one of the most basal Caimanines, even sharing some traits with alligatorids. Etymology The generic name ''Globidentosuchus'' is derived from the Latin roots ''globus'' meaning "sphere" and ''dens'' meaning "tooth", referring to the spherical teeth in the posterior skull, and Greek ''souchos'' meaning "crocodile" after its classification. The species name ''brachyrostris'' is derived from the Greek ''brachys'' meaning "short" and Latin ''rostrum'' meaning "snout" after the truncated and robust rostrum of the species. History and taxonomy ''Globidentosuchus'' was described in 2013 by T. M. Scheyer and colleagues, the ty ...
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Caimaninae
A caiman (also cayman as a variant spelling) is an alligatorid belonging to the subfamily Caimaninae, one of two primary lineages within the Alligatoridae family, the other being alligators. Caimans inhabit Mexico, Central and South America from marshes and swamps to mangrove rivers and lakes. They have scaly skin and live a fairly nocturnal existence. They are relatively small-sized crocodilians with an average maximum weight of depending on species, with the exception of the black caiman (''Melanosuchus niger''), which can grow more than in length and weigh in excess of 1,000 kg (2,200 Ib). The black caiman is the largest caiman species in the world and is found in the slow-moving rivers and lakes that surround the Amazon basin. The smallest species is the Cuvier's dwarf caiman (''Paleosuchus palpebrosus''), which grows to long. There are six different species of caiman found throughout the watery jungle habitats of Central and Southern America. The average length for most o ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Phylogenetic Analysis
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does n ...
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