Eryopoidea
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Eryopoidea
Eryopoidea are a taxon of late Carboniferous and Permian temnospondyli amphibians, known from North America and Europe. Carroll includes no fewer than ten families, but Yates and Warren replace this with a cladistic approach and include three families, the Eryopidae, Parioxyidae and Zatrachydidae. They define the Eryopoidea as all Euskelia in which the choana are relatively rounded and the iliac blade The crest of the ilium (or iliac crest) is the superior border of the wing of ilium and the superiolateral margin of the greater pelvis. Structure The iliac crest stretches posteriorly from the anterior superior iliac spine (ASIS) to the posteri ... vertical. A similar definition but without the Euskelia is provided by Laurin and Steyer. References External linksPalaeos
– Mikko's Phylogeny Archive {{Taxonbar, fro ...
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Temnospondyli
Temnospondyli (from Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') is a diverse order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carboniferous, Permian, and Triassic periods. A few species continued into the Jurassic and Cretaceous periods. Fossils have been found on every continent. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis, and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are considered amphibians, many had characteristics, such as scales and armour-like bon ...
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Eryopidae
Eryopidae were a group of medium to large amphibious temnospondyli, known from North America and Europe. They are defined as all eryopoids with interpterygoid vacuities (spaces in the interpterygoid bone) that are rounded at the front; and large external nares (Laurin and Steyer 2000). Not all of the genera previously included in the Eryopidae (Carroll 1988) are retained under the cladistic revisions. Gallery File:Eryops1DB.jpg, '' Eryops megacephalus'', of the late Carboniferous to early Permian of North America File:Onchiodon12DB.jpg, ''Onchiodon'', of the late Carboniferous to early Permian of Europe and North America File:Actinodon frossardi 1DB.jpg, '' Actinodon frossardi'', of the early Permian of France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ... File:Clamorosaur ...
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Eryops Megacephalus
''Eryops'' (; from Greek , , 'drawn-out' + , , 'face', because most of its skull was in front of its eyes) is a genus of extinct, amphibious temnospondyls. It contains the single species , the fossils of which are found mainly in early Permian (about 295 million years ago) rocks of the Texas Red Beds, located in Archer County, Texas. Fossils have also been found in late Carboniferous period rocks from New Mexico. Several complete skeletons of ''Eryops'' have been found in lower Permian rocks, but skull bones and teeth are its most common fossils. Description ''Eryops'' averaged a little over long and could grow up to , making them among the largest land animals of their time. Adults weighed between . The skull was proportionately large, being broad and flat and reaching lengths of . It had an enormous mouth with many curved teeth like the frog. Its teeth had enamel with a folded pattern, leading to its early classification as a "labyrinthodont" ("maze toothed"). The sha ...
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Parioxyidae
''Parioxys'' is an extinct genus of temnospondyl amphibian from the Early Permian of Texas. History of study The type species, ''Parioxys ferricolus'', was named in 1878 by American paleontologist Edward Drinker Cope based on two badly preserved skulls that were collected from the early Permian Texas red-beds. Egyptian paleontologist Youssef S. Moustafa, described new material of ''P. ferricolus'' from other localities in Texas. This taxon was recently redescribed by Schoch & Sues (2022). Moustafa also described another species, ''P. romeri'', on the basis of an isolated humerus, but this was regarded as being indeterminate. A second definitive species, ''P. bolli'', was described by Canadian paleontologist Robert Carroll in 1964. This taxon is only known from postcranial material. Relationships ''Parioxys'' was historically considered to be closely related to eryopoids, more specifically the well-known '' Eryops megalocephalus'', which was collected from the same locality ...
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Zatrachydidae
Zatracheidae (sometimes mistakenly spelled Zatrachydidae or Zatrachysdidae) is a family of Late Carboniferous and Early Permian temnospondyl amphibians known from North America and Europe. Zatracheidids are distinguished by lateral (sideways) bony protuberances of the quadratojugal bone of the skull, and a large opening in the snout called the internarial fontanelle (sometimes the internarial fenestra) that is bordered by enlarged premaxillae. The skull is flattened, with small orbits or eye sockets set far back. The opening in the snout may have housed a gland for producing a sticky substance so that prey would adhere to the tongue. If so, this indicates that these animals spent a large part of their time on land. History of study There are three genera of zatracheidids: '' Acanthostomatops'' Kuhn, 1961; '' Dasyceps'' Huxley 1859; and '' Zatrachys'' Cope, 1878. Only ''Dascyeps'' is represented by a species in addition to the type species. The name Zatracheidae was first coin ...
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Euskelia
Euskelia is a clade of extinct Temnospondyl amphibians. ''Euskelia'' is a stem-based taxon including all temnospondyls more closely related to ''Eryops'' than to ''Parotosuchus ''Parotosuchus'' is an extinct genus of capitosaurian temnospondyl amphibians within the family Mastodonsauridae. Fossils are known from the Early Triassic of Europe, Africa, Australia, and Antarctica. It was about long and likely lived in aqua ...''. References Carboniferous temnospondyls Permian temnospondyls Triassic temnospondyls Pennsylvanian first appearances Early Triassic extinctions {{temnospondyli-stub es:Euskelia ...
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Pennsylvanian (geology)
The Pennsylvanian ( , also known as Upper Carboniferous or Late Carboniferous) is, in the International Commission on Stratigraphy, ICS geologic timescale, the younger of two period (geology), subperiods (or upper of two system (stratigraphy), subsystems) of the Carboniferous Period. It lasted from roughly . As with most other geochronology, geochronologic units, the stratum, rock beds that define the Pennsylvanian are well identified, but the exact date of the start and end are uncertain by a few hundred thousand years. The Pennsylvanian is named after the U.S. state of Pennsylvania, where the coal-productive beds of this age are widespread. The division between Pennsylvanian and Mississippian (geology), Mississippian comes from North American stratigraphy. In North America, where the early Carboniferous beds are primarily marine limestones, the Pennsylvanian was in the past treated as a full-fledged geologic period between the Mississippian and the Permian. In parts of Europe, ...
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Permian
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids ( reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their am ...
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Edward Drinker Cope
Edward Drinker Cope (July 28, 1840 – April 12, 1897) was an American zoologist, paleontologist, comparative anatomist, herpetologist, and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested in science; he published his first scientific paper at the age of 19. Though his father tried to raise Cope as a gentleman farmer, he eventually acquiesced to his son's scientific aspirations. Cope married his cousin and had one child; the family moved from Philadelphia to Haddonfield, New Jersey, although Cope would maintain a residence and museum in Philadelphia in his later years. Cope had little formal scientific training, and he eschewed a teaching position for field work. He made regular trips to the American West, prospecting in the 1870s and 1880s, often as a member of United States Geological Survey teams. A personal feud between Cope and paleontologist Othniel Charles Marsh led to a period of intense fossil-finding competition ...
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Choana
The choanae (singular choana), posterior nasal apertures or internal nostrils are two openings found at the back of the nasal passage between the nasal cavity and the throat in tetrapods, including humans and other mammals (as well as crocodilians and most skinks). They are considered one of the most important synapomorphies of tetrapodomorphs, that allowed the passage from water to land. In animals with secondary palates, they allow breathing when the mouth is closed. Janvier, Philippe (2004) "Wandering nostrils". ''Nature'', 432 (7013): 23–24. In tetrapods without secondary palates their function relates primarily to olfaction (sense of smell). The choanae are separated in two by the vomer. Boundaries A choana is the opening between the nasal cavity and the nasopharynx. It is therefore not a structure but a space bounded as follows: * anteriorly and inferiorly by the horizontal plate of palatine bone, * superiorly and posteriorly by the sphenoid bone * laterally by t ...
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Iliac Blade
The crest of the ilium (or iliac crest) is the superior border of the wing of ilium and the superiolateral margin of the greater pelvis. Structure The iliac crest stretches posteriorly from the anterior superior iliac spine (ASIS) to the posterior superior iliac spine (PSIS). Behind the ASIS, it divides into an outer and inner lip separated by the intermediate zone. The outer lip bulges laterally into the iliac tubercle. Platzer (2004), p 186 Palpable in its entire length, the crest is convex superiorly but is sinuously curved, being concave inward in front, concave outward behind. Palastanga (2006), p 243 It is thinner at the center than at the extremities. Development The iliac crest is derived from endochondral bone. Function To the external lip are attached the ''Tensor fasciae latae'', '' Obliquus externus abdominis'', and ''Latissimus dorsi'', and along its whole length the ''fascia lata''; to the intermediate line, the '' Obliquus internus abdominis''. To the interna ...
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Carboniferous Temnospondyls
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferous'' means "coal-bearing", from the Latin '' carbō'' ("coal") and '' ferō'' ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern 'system' names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession. The Carboniferous is often treated in North America as two geological periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial animal life was well established by the Carboniferous Period. Tetrapods (four limbed vertebrates), which had originated from lobe-finned fish during the preceding Devonian, became pentadactylous in and diversified during the Carboniferous, including early amphibian lineages ...
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