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Enteropneust
The acorn worms or Enteropneusta are a hemichordate class of invertebrates consisting of one order of the same name. The closest non-hemichordate relatives of the Enteropneusta are the echinoderms. There are 111 known species of acorn worm in the world, the main species for research being ''Saccoglossus kowalevskii''. Two families—Harrimaniidae and Ptychoderidae—separated at least 370 million years ago. Until recently, it was thought that all species lived in the sediment on the seabed, subsisting as deposit feeders or suspension feeders. However, the early 21st century has seen the description of a new family, the Torquaratoridae, evidently limited to the deep sea, in which most of the species crawl on the surface of the ocean bottom and alternatively rise into the water column, evidently to drift to new foraging sites. It is assumed that the ancestors of acorn worms used to live in tubes like their relatives Pterobranchia, but that they eventually started to live a safer a ...
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Saccoglossus Kowalevskii
Hemichordata is a phylum which consists of triploblastic, enterocoelomate, and bilaterally symmetrical marine deuterostome animals, generally considered the sister group of the echinoderms. They appear in the Lower or Middle Cambrian and include two main classes: Enteropneusta (acorn worms), and Pterobranchia. A third class, Planctosphaeroidea, is known only from the larva of a single species, ''Planctosphaera pelagica''. The class Graptolithina, formerly considered extinct, is now placed within the pterobranchs, represented by a single living genus ''Rhabdopleura''. Acorn worms are solitary worm-shaped organisms. They generally live in burrows (the earliest secreted tubes) and are deposit feeders, but some species are pharyngeal filter feeders, while the family Torquaratoridae are free living detritivores. Many are well known for their production and accumulation of various halogenated phenols and pyrroles. Pterobranchs are filter-feeders, mostly colonial, living in a colla ...
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Pterobranchia
Pterobranchia is a class of small worm-shaped animals. They belong to the Hemichordata, and live in secreted tubes on the ocean floor. Pterobranchia feed by filtering plankton out of the water with the help of cilia attached to tentacles. There are about 25 known living pterobranch species in three genera, which are ''Rhabdopleura'', '' Cephalodiscus'', and '' Atubaria''. On the other hand, there are several hundred extinct genera, some of which date from the Cambrian Period. The class Pterobranchia was established by Ray Lankester in 1877. It contained, at that time, the single genus ''Rhabdopleura''. ''Rhabdopleura'' was at first regarded as an aberrant polyzoon, but when the ''Challenger'' report on '' Cephalodiscus'' was published in 1887, it became clear that ''Cephalodiscus'', the second genus now included in the order, had affinities with the Enteropneusta. Electron microscope studies have suggested that pterobranchs belong to the same clade as the extinct grapto ...
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Balanoglossus 2
''Balanoglossus'' is a genus of ocean-dwelling acorn worms (Enteropneusta). It has zoological importance because, being hemichordates, they are an "evolutionary link" between invertebrates and vertebrates. ''Balanoglossus'' is a deuterostome, and resembles the sea squirts (Ascidiacea) in that it possesses branchial openings, or "gill slits". It has a notochord in the upper part of the body and has no nerve chord. It does have a stomochord, however, which is a gut chord within the collar. Their heads may be as small as per 2.5 mm (1/10 in) or as large as 5 mm (1/5 in). Discovery The discovery of gill-slits in this animal by Alexander Kovalevsky (1865) led to the creation of the class Enteropneusta by Karl Gegenbaur (1870). Classification William Bateson (1885) originally included them in phylum Chordata. Hyman (1959), however, placed them near Echinodermata and gave Hemichordata a status of an independent phylum. Habitat Balanoglossus is a tuberculos (bur ...
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Balanoglossus 3
''Balanoglossus'' is a genus of ocean-dwelling acorn worms (Enteropneusta). It has zoological importance because, being hemichordates, they are an "evolutionary link" between invertebrates and vertebrates. ''Balanoglossus'' is a deuterostome, and resembles the sea squirts (Ascidiacea) in that it possesses branchial openings, or "gill slits". It has a notochord in the upper part of the body and has no nerve chord. It does have a stomochord, however, which is a gut chord within the collar. Their heads may be as small as per 2.5 mm (1/10 in) or as large as 5 mm (1/5 in). Discovery The discovery of gill-slits in this animal by Alexander Kovalevsky (1865) led to the creation of the class Enteropneusta by Karl Gegenbaur (1870). Classification William Bateson (1885) originally included them in phylum Chordata. Hyman (1959), however, placed them near Echinodermata and gave Hemichordata a status of an independent phylum. Habitat Balanoglossus is a tuberculos (bur ...
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Balanoglossus Gigas
''Balanoglossus'' is a genus of ocean-dwelling acorn worms (Enteropneusta). It has zoological importance because, being hemichordates, they are an "evolutionary link" between invertebrates and vertebrates. ''Balanoglossus'' is a deuterostome, and resembles the sea squirts (Ascidiacea) in that it possesses branchial openings, or "gill slits". It has a notochord in the upper part of the body and has no nerve chord. It does have a stomochord, however, which is a gut chord within the collar. Their heads may be as small as per 2.5 mm (1/10 in) or as large as 5 mm (1/5 in). Discovery The discovery of gill-slits in this animal by Alexander Kovalevsky (1865) led to the creation of the class Enteropneusta by Karl Gegenbaur (1870). Classification William Bateson (1885) originally included them in phylum Chordata. Hyman (1959), however, placed them near Echinodermata and gave Hemichordata a status of an independent phylum. Habitat Balanoglossus is a tuberculos (bur ...
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Spartobranchus Tenuis
Spartobranchus tenuis is an extinct species of acorn worms (Enteropneusta). It existed in the Middle Cambrian (505 million years ago). Petrified mark animals were found in British Columbia, Canada in the Burgess Shale formation. It is similar to the modern representatives of the family Harrimaniidae, distinguished by branching fiber tubes. It is a believed predecessor of Pterobranchia Pterobranchia is a class of small worm-shaped animals. They belong to the Hemichordata, and live in secreted tubes on the ocean floor. Pterobranchia feed by filtering plankton out of the water with the help of cilia attached to tentacles. The ..., but this species is intermediate between these two classes. Studies show that these tubes were lost in the line leading to modern acorn worm, but remained in the extinct graptolites and saving still perystozyabernyh. Description Detailed analysis shows that ''Spartobranchus tenuis'' had a flexible body consisting of short proboscis, collar and narr ...
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Torquaratoridae
Torquaratoridae (Latin for "neck plow") is a family of acorn worms ( Hemichordata) that lives in deep waters between 350 to 4000 meters (the species Tergivelum baldwinae has been found 4100 meters below the surface). They can grow up to three feet in length and have semitransparent gelatinous bodies, often brightly colored. Cilia on their underside are used to glide over the ocean floor at about three inches per hour, while detritus is sucked into their gut, leaving behind a constant trail of feces. When deciding to move to new feeding locations, they empty their gut and drift over the bottom, aided by an excreted balloon of mucus, before they let themselves down somewhere else. One species (''Coleodesmium karaensis'') has been shown to care for the offspring by bearing about a dozen embryos surrounded by a thin membrane in shallow depressions on the surface of the mother's pharyngeal region. The proboscis skeleton is reduced to a small medial plate in one genus, while it is ab ...
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Peristalsis
Peristalsis ( , ) is a radially symmetrical contraction and relaxation of muscles that propagate in a wave down a tube, in an anterograde direction. Peristalsis is progression of coordinated contraction of involuntary circular muscles, which is preceded by a simultaneous contraction of the longitudinal muscle and relaxation of the circular muscle in the lining of the gut. In much of a digestive tract such as the human gastrointestinal tract, smooth muscle tissue contracts in sequence to produce a peristaltic wave, which propels a ball of food (called a bolus before being transformed into chyme in the stomach) along the tract. The peristaltic movement comprises relaxation of circular smooth muscles, then their contraction behind the chewed material to keep it from moving backward, then longitudinal contraction to push it forward. Earthworms use a similar mechanism to drive their locomotion, and some modern machinery imitate this design. The word comes from New Latin and is ...
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Iodoform
Iodoform (also known as triiodomethane and, inaccurately, as carbon triiodide) is the organoiodine compound with the chemical formula C H I3. A pale yellow, crystalline, volatile substance, it has a penetrating and distinctive odor (in older chemistry texts, the smell is sometimes referred to as that of hospitals, where the compound is still commonly used) and, analogous to chloroform, sweetish taste. It is occasionally used as a disinfectant. Structure The molecule adopts tetrahedral molecular geometry with C3v symmetry. Synthesis and reactions The synthesis of iodoform was first described by Georges-Simon Serullas in 1822, by reactions of iodine vapour with steam over red-hot coals, and also by reaction of potassium with ethanolic iodine in the presence of water; and at much the same time independently by John Thomas Cooper. It is synthesized in the haloform reaction by the reaction of iodine and sodium hydroxide with any one of these four kinds of organic compounds: a methyl ...
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Cilium
The cilium, plural cilia (), is a membrane-bound organelle found on most types of eukaryotic cell, and certain microorganisms known as ciliates. Cilia are absent in bacteria and archaea. The cilium has the shape of a slender threadlike projection that extends from the surface of the much larger cell body. Eukaryotic flagella found on sperm cells and many protozoans have a similar structure to motile cilia that enables swimming through liquids; they are longer than cilia and have a different undulating motion. There are two major classes of cilia: ''motile'' and ''non-motile'' cilia, each with a subtype, giving four types in all. A cell will typically have one primary cilium or many motile cilia. The structure of the cilium core called the axoneme determines the cilium class. Most motile cilia have a central pair of single microtubules surrounded by nine pairs of double microtubules called a 9+2 axoneme. Most non-motile cilia have a 9+0 axoneme that lacks the central pair o ...
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Mouth
In animal anatomy, the mouth, also known as the oral cavity, or in Latin cavum oris, is the opening through which many animals take in food and issue vocal sounds. It is also the cavity lying at the upper end of the alimentary canal, bounded on the outside by the lips and inside by the pharynx. In tetrapods, it contains the tongue and, except for some like birds, teeth. This cavity is also known as the buccal cavity, from the Latin ''bucca'' ("cheek"). Some animal phyla, including arthropods, molluscs and chordates, have a complete digestive system, with a mouth at one end and an anus at the other. Which end forms first in ontogeny is a criterion used to classify bilaterian animals into protostomes and deuterostomes. Development In the first multicellular animals, there was probably no mouth or gut and food particles were engulfed by the cells on the exterior surface by a process known as endocytosis. The particles became enclosed in vacuoles into which enzymes were secr ...
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Diverticulum
In medicine or biology, a diverticulum is an outpouching of a hollow (or a fluid-filled) structure in the body. Depending upon which layers of the structure are involved, diverticula are described as being either true or false. In medicine, the term usually implies the structure is not normally present, but in embryology, the term is used for some normal structures arising from others, as for instance the thyroid diverticulum, which arises from the tongue. The word comes from Latin ''dīverticulum'', "bypath" or "byway". Classification Diverticula are described as being true or false depending upon the layers involved: *False diverticula (also known as "pseudodiverticula") do not involve muscular layers or adventitia. False diverticula, in the gastrointestinal tract for instance, involve only the submucosa and mucosa. *True diverticula involve all layers of the structure, including muscularis propria and adventitia, such as Meckel's diverticulum. Embryology *The kidneys are ...
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