Eburoceras
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Eburoceras
''Eburoceras'' is an early nautiloid cephalopod from the Upper Yenchau and Wanwankau, Upper Cambrian Trempealeauan of China, assigned to the Ellesmeroceratidae. The shell of ''Eburoceras'' is long and narrow, curved in a broad arc of near constant curvature, interpreted as endogastric, assuming the small siphuncle on the inner margin to be ventral. Known for sure dorsal siphuncles do not appear until later in the Early Ordovician with the Trocholitidae, which if so would make it (''Eburoceras'') exogastric. ''Eburoceras'' is the most strongly curved of the Cambrian ellesmeroceratids. Eburoceras is the earliest ellesmeroceratid except for the short lived '' Hunyuanocers'', from the lower part of the Upper Yenchou, that proceeded it. ''Eburoceras'' died out at the end of Wanwankau time, along with the other Late Cambrian ellesmeroceratid genera, except for ''Ectenolites ''Ectenolites'' is a genus of small, slender, cylindrical Ellesmeroceratids that resemble '' Ellesmerocera ...
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Ellesmeroceratidae
The Ellesmeroceratidae constitute a family within the cephalopod order Ellesmerocerida. They lived from the Upper Cambrian to the Lower Ordovician. They are characterized by straight and endogastric shells, often laterally compressed, so the dorso-ventral dimension is slightly greater than the lateral, with close spaced sutures having shallow lateral lobes and a generally large tubular ventro-marginal siphuncle with concave segments and irregularly spaced diaphragms. Connecting rings are thick and layered, externally straight but thickening inwardly with the maximum near the middle of the segment so as to leave concave depressions on internal siphuncle molds. Septal necks are typically orthochoanitic but vary in length from almost absent (achoanitic) to reaching halfway to the previous septum (hemichoanitic) and may even slope inwardly (loxochoanitic). Evolution and phylogeny The Ellesmeroceratidae have their derivation in the Plectronoceratidae, order Plectronocerida, in Tr ...
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Cambrian
The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Ca ...
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Trempealeauan
The Trempealeauan is the upper or latest stage of the Upper or Late Cambrian in North America, spanning about 4 million years from about 492.5 to 488.3 m.y.a., equivalent to the Fengshanian of China. The name comes from the Trempealeau Formation, named for the town of Trempealeau in western Wisconsin, located on the Mississippi River. The Trempealeauan follows, or overlies, the Franconian, which is the middle stage of the Upper Cambrian in North America and is followed by the Gasconadian in the Lower Ordovician. Together with the Dresbachian The Dresbachian is a Maentwrogian regional stage of North America, lasting from 501 to 497 million years ago. It is part of the Upper Cambrian and is defined by four trilobite zones. It overlaps with the International Commission on Stratigraphy, IC ... at the bottom, the Trempealeauan and Franconian make up the Croixan Series. References * Chen Jun-yuan & Teichert C, 1983; Cambrian Cephalopods, Geology Vol 11, pp647–650, Nov 1983 * Flowe ...
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Animalia
Animals are multicellular, eukaryotic organisms in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a bilaterally symmetric body plan. The Bilateria include the protostomes, containing animals such as nematodes, arthropods, flatworms, annelids and molluscs, and the deuterostomes, containing the echinode ...
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Cephalopoda
A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by ''Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been identified. Tw ...
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Nautiloidea
Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and ''Allonautilus''. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms ( orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day. In a broad sense, "nautiloid" refers to a major cephalopod subclass or collection of subclasses (Nautiloidea ''sensu lato''). Nautiloids are typically considered one of three main groups of cephalopods, along with the extinct ammonoids (ammonites) and living coleoids (such as squid, octopus, and kin). While ammonoids and coleoids are monophyletic clades with exclusive ancestor-descendant rela ...
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Ellesmerocerida
The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician. Morphology The Ellesmerocerida are characterized by shells that are typically small, some even tiny, with close-spaced septa and relatively large ventral siphuncles. In some genera (e.g. ''Paleoceras''), the septa are uniformly spaced. Shells of ellesmerocerids are typically smooth and compressed and vary in form. They may be breviconic (short) or longiconic (elongate), straight (orthoconic) or curved (cyrtoconic). Cyrtoconic forms are usually endogastric, with longitudinally convex ventral margins. The apeces of straight forms typically have an endogastric curvature. Some may have grown to as much as 15 cm. Siphuncle segments are tubular or concave. Septal necks are short. Connecting rings which may appear layered are thick and typically wedge shaped with their maximum width at or near where t ...
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Trocholitidae
The Trocholitidae are Tarphycerida with whorls in close contact as with the Tarphyceratidae, but in which the siphuncle, similar in structure, becomes dorsal. The Trocholitidae are derived from the Tarphyceratidae, perhaps from different tarphyceratids. Members of the Trocholitidae vary in whorl section and siphuncle position. Most are tightly coiled with the dorsum impressed to some degree. As exceptions ''Arkoceras'' and ''Wichitoceras'' have their whorls touching but without a dorsal impression. '' Discoceras'' and '' Curtoceras'' have somewhat subquadrate whorl sections and are moderately impressed. In ''Graftonoceras'' and ''Jasperoceras'' the venter is rounded and continues smoothly through the flanks to a broad dorsum which in each is moderately impressed. The siphuncle in ''Graftonoceras'' is on the dorsal margin, in ''Jasperoceras'' it is between the center and dorsum. Whorls in T''rocholites'' have a wide cross section and low profile, in ''Wichitoceras'' they are lat ...
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Ectenolites
''Ectenolites'' is a genus of small, slender, cylindrical Ellesmeroceratids that resemble '' Ellesmeroceras'' but are smaller and proportionally narrower. Septa, as typical for ellesmerocerids, are close spaced with shallow lobes on either flank. The body chamber is proportionally long, the shell itself slightly compressed. The dorsal side at the beginning of the shell, opposite the aperture and body chamber, is strongly convex so to produce a sense of endogastric curvature with the apex and siphuncle The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and ... aligned.Flower R.H.1964, The Nautiloid Order Ellesmeroceratida (Cephalopoda). Memoir 12, New Mexico Bureau of Mines and Mineral Resources, Socorro, NM The siphuncle in ''Ectenolites'' lies along the ventral side; is tubular, composed ...
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Clarkoceras
''Clarkoceras'' is a genus of breviconic ellesmerocerid cephalopods, one of only two genera known to have crossed from the Late Cambrian, Trempealeauan, into the Early Ordovician, Gasconadian. (Flower 1964, Teichert 1988); the other being '' Ectenolites''. Description Clarkoceras has a rapidly expanding, laterally compressed, relatively short, endogastrically cyrtoconic shell; The upper or dorsal side is more strongly convex longitudinally than the lower or ventral side is concave. Sutures are essentially straight and close spaced indicating very short camerae (chambers). The siphuncle is relatively large, 0.3 the dorsoventral dimension and is ventral, although not necessarily marginal. (Flower 1964, Furnish and Glenister 1964). Septal necks are straight, reaching about half way back to the previous septum and the connecting rings are thick and layered Taxonomy ''Clarkoceras'' was first thought by Clarke to be a pilocerid, who in 1897 give it the name ''Piloceras newton-winche ...
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Ordovician
The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. The Ordovician, named after the Celtic Britons, Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same Rock (geology), rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed Stratum, strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Union of Geological Sciences, Intern ...
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