Dryolestida
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Dryolestida
Dryolestida is an extinct order of mammals, primarily and possibly exclusively known from the Jurassic and Cretaceous. They are considered members of the clade Cladotheria, close to the ancestry of therian mammals. It is also believed that they developed a fully mammalian jaw and also had the three middle ear bones. Most members of the group, as with most Mesozoic mammals, are only known from fragmentary tooth and jaw remains. The taxonomic composition of the group is contested. Aside from the uncontroversial Dryolestidae and the possibly paraphyletic Paurodontidae, which were small insectivores, known from the Middle Jurassic to Early Cretaceous of Laurasia, the Meridiolestida, a diverse group of mammals including both small insectivores and mid-large sized herbivores known from the Late Cretaceous to Miocene of South America and possibly Antarctica, are also often included. However, in many phylogenetic analyses, Meridiolestida are recovered as an unrelated group of cladotheri ...
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Brandoniidae
Dryolestidae is an extinct family of Mesozoic mammals, known from the Middle Jurassic to the Early Cretaceous of the North Hemisphere. The oldest known member, '' Anthracolestes'', is known from the Middle Jurassic Itat Formation of Western Siberia, but most other representatives are known from the Late Jurassic of North America and the Late Jurassic and Early Cretaceous of Europe. Most members are only known from isolated teeth and jaw fragments. Like many other groups of early mammals, they are though to have been insectivores. They are generally classified in Cladotheria, meaning that they are considered to be more closely related to marsupials and placentals than to monotremes. They are placed as part of the broader Dryolestida, which also includes the (possibly paraphyletic) Paurodontidae, and also sometimes the South American-Antarctic Meridiolestida, which are often considered unrelated cladotherians. Dyolestidae is not based on a phylogenetic definition, but instead on ...
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Dryolestidae
Dryolestidae is an extinct family of Mesozoic mammals, known from the Middle Jurassic to the Early Cretaceous of the North Hemisphere. The oldest known member, '' Anthracolestes'', is known from the Middle Jurassic Itat Formation of Western Siberia, but most other representatives are known from the Late Jurassic of North America and the Late Jurassic and Early Cretaceous of Europe. Most members are only known from isolated teeth and jaw fragments. Like many other groups of early mammals, they are though to have been insectivores. They are generally classified in Cladotheria, meaning that they are considered to be more closely related to marsupials and placentals than to monotremes. They are placed as part of the broader Dryolestida, which also includes the (possibly paraphyletic) Paurodontidae, and also sometimes the South American-Antarctic Meridiolestida, which are often considered unrelated cladotherians. Dyolestidae is not based on a phylogenetic definition, but instead on ...
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Henkelotherium
''Henkelotherium'' is an extinct genus of dryolestidan mammal from the Late Jurassic (Kimmeridgian) Camadas de Guimarota, in Portugal. Unlike many other Jurassic mammals, it is known from a largely complete skeleton, and is thought to have had an arboreal lifestyle. Description The skull of ''Henkelotherium'' is long, and presacral body length is . This suggest a weight of about . Paleobiology Primitive characters of ''Henkelotherium'' (e.g. asymmetric condyles of the femur) indicate that this species had a mode of locomotion similar to tree shrews and opossums. The small size of ''Henkelotherium'' and elongated tail made it suited to an arboreal lifestyle and capable of climbing trees, a notion supported by the paleoecological reconstruction of the Guimarota ecosystem indicating a densely vegetated environment. Taxonomy In cladistic analyses, ''Henkelotherium'' has been considered closely related to Dryolestidae, either as a part of that group, or as closely related b ...
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Pantotheria
Pantotheria is an abandoned taxon of Mesozoic mammals. This group is now considered an informal "wastebasket" taxon and has been replaced by Dryolestida as well as other groups. It is sometimes treated as an infraclass and older books refer to it as being related to symmetrodonts. One classification makes it an infraclass with a single order, Eupantotheria. Taxonomy List of mammals that were at one time included in the group PantotheriaMikko's Phylogeny Archiv *Genus †'' Tribactonodon, Tribactonodon bonfieldi'' Sigogneau-Russell, Hooker & Ensom 2001 *Genus †'' Paraungulatum rectangularis'' Bonaparte 1999 *Genus †'' Argaliatherium robustum '' Cifelli & Davis 2015 *Genus †'' Carinalestes murensis'' Cifelli & Davis 2015 *Genus †'' Hypomylos'' Sigogneau-Russell 1992 *Family † Picopsidae Fox 1980 **Genus †'' Picopsis pattersoni'' Fox 1980 **Genus †'' Tirotherium aptum'' Montellano-Ballesteros & Fox 2015 *Family † Casamiqueliidae Bonaparte 1999 **Genus †'' Casa ...
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Cladotheria
Cladotheria is a clade (sometimes ranked as a legion) of mammals. It contains modern therian mammals (marsupials and placentals) and several extinct groups, such as the dryolestoids, amphitheriids and peramurids. The clade was named in 1975 by Malcolm McKenna. In 2002, it was defined as a node-based taxon containing "the common ancestor of dryolestids and living therians, plus all its descendants". A different, stem-based definition was given in 2013, in which Cladotheria contains all taxa that are closer to ''Mus musculus'' (the house mouse) than to the "symmetrodont" '' Spalacotherium tricuspidens''. Description Early cladotherians can be distinguished from other mammals by a number of derived traits (apomorphies). Their teeth differed from those of the "symmetrodonts" by the evolution of a talonid shelf ( hypoflexid) on the lower molars, which occluded with the paracone of the corresponding upper molars. A true talonid basin, allowing for the crushing and grinding of f ...
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Eupantotheria
Pantotheria is an abandoned taxon of Mesozoic mammals. This group is now considered an informal "wastebasket" taxon and has been replaced by Dryolestida as well as other groups. It is sometimes treated as an infraclass and older books refer to it as being related to symmetrodonts. One classification makes it an infraclass with a single order, Eupantotheria. Taxonomy List of mammals that were at one time included in the group PantotheriaMikko's Phylogeny Archiv *Genus †'' Tribactonodon, Tribactonodon bonfieldi'' Sigogneau-Russell, Hooker & Ensom 2001 *Genus †'' Paraungulatum rectangularis'' Bonaparte 1999 *Genus †'' Argaliatherium robustum '' Cifelli & Davis 2015 *Genus †'' Carinalestes murensis'' Cifelli & Davis 2015 *Genus †'' Hypomylos'' Sigogneau-Russell 1992 *Family † Picopsidae Fox 1980 **Genus †'' Picopsis pattersoni'' Fox 1980 **Genus †'' Tirotherium aptum'' Montellano-Ballesteros & Fox 2015 *Family † Casamiqueliidae Bonaparte 1999 **Genus †'' Casa ...
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Meridiolestida
Meridiolestida is an extinct clade of mammals known from the Cretaceous and Cenozoic of South America and possibly Antarctica. They represented the dominant group of mammals in South America during the Late Cretaceous. Meridiolestidans were morphologically diverse, containing both small insectivores such as the "sabretooth-squirrel" '' Cronopio,supplementary information
'' as well as the clade Mesungulatoidea/Mesungulatomorpha, which ranged in size from the shrew-sized '''' to the dog-sized ''

Paurodontidae
Paurodontidae is a family of Late Jurassic to Early Cretaceous mammals in the order Dryolestida. Remains of paurodontids have been found in the United States, Britain, Portugal, and Tanzania. The group likely represents a paraphyletic group of basal non dryolestid dryolestidans. ''Paurodon'' has been suggested to have been a specilast feeder on earthworms due to the morphology of its teeth closely resembling that of the golden mole genus ''Amblysomus ''Amblysomus'' (also narrow-headed golden mole or South African golden mole) is a genus of the golden mole family, Chrysochloridae, comprising five species of the small, insect-eating, burrowing mammals endemic to Southern Africa. All five spec ...''.A.O. Averianov and T. Martin (2015). "Ontogeny and taxonomy of Paurodon valens (Mammalia, Cladotheria) from the Upper Jurassic Morrison Formation of USA" (PDF). Proceedings of the Zoological Institute of the Russian Academy of Sciences 319 (3): 326–340. References Dryolesti ...
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Dryolestes
''Dryolestes'' is an extinct genus of Late Jurassic mammal from the Morrison Formation and the Alcobaça Formation The Alcobaça Formation, previously known as the Guimarota Formation and also known as the Consolação Unit, is a geological formation in Portugal. It dates back to the Kimmeridgian stage of the Late Jurassic. It is an important source of inform ... of Portugal. The type species ''Dryolestes priscus'' is present in stratigraphic zones 2, 5, and 6.Foster, J. (2007). "Appendix." ''Jurassic West: The Dinosaurs of the Morrison Formation and Their World''. Indiana University Press. pp. 327-329. See also * Prehistoric mammal ** List of prehistoric mammals * Paleobiota of the Morrison Formation References Further reading * Foster, J. (2007). Jurassic West: The Dinosaurs of the Morrison Formation and Their World. Indiana University Press. 389pp. Dryolestida Kimmeridgian genus first appearances Tithonian genus extinctions Jurassic mammals of Europe Ju ...
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Jurassic
The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period of the Mesozoic, Mesozoic Era and is named after the Jura Mountains, where limestone strata from the period were first identified. The start of the Jurassic was marked by the major Triassic–Jurassic extinction event, associated with the eruption of the Central Atlantic magmatic province, Central Atlantic Magmatic Province. The beginning of the Toarcian Stage started around 183 million years ago and is marked by an extinction event associated with widespread Anoxic event, oceanic anoxia, ocean acidification, and elevated temperatures likely caused by the eruption of the Karoo-Ferrar, Karoo-Ferrar large igneous provinces. The end of the Jurassic, however, has no clear boundary with the Cretaceous and i ...
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Mesungulatids
Mesungulatidae is an extinct clade of meridiolestidan dryolestoid mammals from the Late Cretaceous of South America and possibly other Gondwannan landmasses. They are particularly notable for their ecological speciation and large size. Characteristics Most mesungulatids are generally large animals, making them inherently distinctive from other groups. Specific synapomorphies include a strong precingulum and postcingulum on the upper molars - which are extended lingually but do not meet around the paracone - three cusps on the lower stylar shelf, an absent metacone and rectangular lower molars. They are thought to have had a somewhat transverse mastication, like docodonts and modern ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, ...s. Compared to other dryolestoids thei ...
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Bathonian
In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Stratigraphic definitions The Bathonian Stage takes its name from Bath, a spa town in England built on Jurassic limestone (the Latinized form of the town name is ''Bathonium''). The name was introduced in scientific literature by Belgian geologist d'Omalius d'Halloy in 1843. The original type locality was located near Bath. The French palaeontologist Alcide d'Orbigny was in 1852 the first to define the exact length of the stage. The base of the Bathonian is at the first appearance of ammonite species '' Parkinsonia (Gonolkites) convergens'' in the stratigraphic column. The global reference profile for the base of the Bathonian (a GSSP) was ratified as Ravin du Bès, Bas-Auran area, Alpes de Haute Provence, France in 2009. The top of the Bat ...
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