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Dromaeosaurinae
Dromaeosaurinae is a subfamily of the theropod group Dromaeosauridae. The earliest dromaeosaurine is ''Utahraptor'', dating back to the Early Cretaceous period in North America, however, some isolated teeth seems to represent an indeterminate species of dromaeosaurine, coming from the Late Jurassic period in Africa. If true, this will push their range to the Jurassic period, instead of the Cretaceous, as in most dromaeosaurs. Most dromaeosaurs are small carnivores, however, dromaeosaurines are represented by some of the largest species (''Achillobator'', ''Dakotaraptor'', ''Utahraptor''). So far, ''Dakotaraptor'' is the only dromaeosaurine with evidence of quill knobs, indicating a plumage; based on this, other members likely had them. Paleobiogeography Most dromaeosaurines lived in what is now Asia, North America and possibly Denmark during the Cretaceous period, from the Berriasian to the Maastrichtian stages. However, isolated teeth that may belong to African dromaeosaurines h ...
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Dromaeosauridae
Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning 'runner', and ('), meaning 'lizard'. In informal usage, they are often called raptors (after ''Velociraptor''), a term popularized by the film '' Jurassic Park''; a few types include the term "raptor" directly in their name and have come to emphasize their bird-like appearance and speculated bird-like behavior. Dromaeosaurid fossils have been found across the globe in North America, Europe, Africa, Asia, South America and Antarctica, with some fossils giving credence to the possibility that they inhabited Australia as well. They first appeared in the mid-Jurassic Period (late Bathonian stage, about 167 million years ago) and survived until the end of the Cretaceous (Maastrichtian stage, 66 ma), existing until the Cretaceous–Paleogene ex ...
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Dromaeosaurus Albertensis
''Dromaeosaurus'' (, "running lizard") is a genus of dromaeosaurid theropod dinosaur which lived during the Late Cretaceous period (middle late Campanian and Maastrichtian), sometime between 80 and 69 million years ago, in Alberta, Canada and the western United States. The type species is ''Dromaeosaurus albertensis'', which was described by William Diller Matthew and Barnum Brown in 1922. Its fossils were unearthed in the Dinosaur Park Formation. Teeth attributed to this genus have been found in the Prince Creek Formation. ''Dromaeosaurus'' is the type genus of both Dromaeosauridae and Dromaeosaurinae, which include many genera with similar characteristics to ''Dromaeosaurus'' such as possibly its closest relative ''Dakotaraptor''. ''Dromaeosaurus'' was heavily built, more so than other dromaeosaurs that are similar in size, like ''Velociraptor''. Discovery and naming Despite receiving widespread attention in popular books on dinosaurs, and the usage of a complete mounted skelet ...
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Dromaeosaurus
''Dromaeosaurus'' (, "running lizard") is a genus of dromaeosaurid theropod dinosaur which lived during the Late Cretaceous period (middle late Campanian and Maastrichtian), sometime between 80 and 69 million years ago, in Alberta, Canada and the western United States. The type species is ''Dromaeosaurus albertensis'', which was described by William Diller Matthew and Barnum Brown in 1922. Its fossils were unearthed in the Dinosaur Park Formation. Teeth attributed to this genus have been found in the Prince Creek Formation. ''Dromaeosaurus'' is the type genus of both Dromaeosauridae and Dromaeosaurinae, which include many genera with similar characteristics to ''Dromaeosaurus'' such as possibly its closest relative '' Dakotaraptor''. ''Dromaeosaurus'' was heavily built, more so than other dromaeosaurs that are similar in size, like ''Velociraptor''. Discovery and naming Despite receiving widespread attention in popular books on dinosaurs, and the usage of a complete mounted skel ...
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Deinonychus
''Deinonychus'' ( ; ) is a genus of dromaeosaurid theropod dinosaur with one described species, ''Deinonychus antirrhopus''. This species, which could grow up to long, lived during the early Cretaceous Period, about 115–108 million years ago (from the mid-Aptian to early Albian stages). Fossils have been recovered from the U.S. states of Montana, Utah, Wyoming, and Oklahoma, in rocks of the Cloverly Formation, Cedar Mountain Formation and Antlers Formation, though teeth that may belong to ''Deinonychus'' have been found much farther east in Maryland. Paleontologist John Ostrom's study of ''Deinonychus'' in the late 1960s revolutionized the way scientists thought about dinosaurs, leading to the " dinosaur renaissance" and igniting the debate on whether dinosaurs were warm-blooded or cold-blooded. Before this, the popular conception of dinosaurs had been one of plodding, reptilian giants. Ostrom noted the small body, sleek, horizontal posture, ratite-like spine, and esp ...
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Velociraptorinae
Velociraptorinae is a subfamily of the theropod group Dromaeosauridae. The earliest velociraptorines are probably ''Nuthetes'' from the United Kingdom, and possibly ''Deinonychus'' from North America. However, several indeterminate velociraptorines have also been discovered, dating to the Kimmeridgian stage, in the Late Jurassic Period. These fossils were discovered in the Langenberg quarry, Oker near Goslar, Germany.van der Lubbe, T., Richter, U. and Knotschke, N. (2009).Velociraptorine dromaeosaurid teeth from the Kimmeridgian (Late Jurassic) of Germany" ''Acta Palaeontologica Polonica'', 54(3): 401-408. Description While most velociraptorines were generally small animals, at least one species may have achieved gigantic sizes comparable to those found among the dromaeosaurines. So far, this unnamed giant velociraptorine is known only from isolated teeth found on the Isle of Wight, England. The teeth belong to an animal the size of dromaeosaurines of the genus ''Utahraptor'', bu ...
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Utahraptor
''Utahraptor'' (meaning "Utah's thief") is a genus of large dromaeosaurid dinosaur that lived in North America during the Early Cretaceous period. It was a heavy-built, ground-dwelling, bipedal carnivore. It contains a single species, ''Utahraptor ostrommaysi'', which is one of the largest-known members of the family Dromaeosauridae, measuring long and weighing . Discovery and naming The first specimens of ''Utahraptor'' were found in 1975 by Jim Jensen in the Dalton Wells Quarry in east-central Utah, near the town of Moab, but did not receive much attention. After a find of a large foot-claw by Carl Limone in October 1991, James Kirkland, Robert Gaston, and Donald Burge uncovered further remains of ''Utahraptor'' in 1991 in the Gaston Quarry in Grand County, Utah, within the Yellow Cat and Poison Strip members of the Cedar Mountain Formation. The holotype of ''Utahraptor'', CEUM 184v.86 consists of a second pedal ungual, with potentially assigned elements from other specim ...
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Achillobator
''Achillobator'' ( ; meaning "Achilles hero") is a genus of large dromaeosaurid dinosaurs that lived in Asia during the Late Cretaceous period about 96 million to 89 million years ago in what is now the Bayan Shireh Formation. The genus is currently monotypic, only including the type species ''A. giganticus''. The first remains were found in 1989 during a Mongolian-Russian field expedition in Mongolia and later described in 1999. Remains at the type locality of ''Achillobator'' may represent additional specimens. It represents the first and largest dromaeosaurid known from the Bayan Shireh Formation. It was a large, heavy-built, ground-dwelling, bipedal carnivore that would have been an active feathered predator hunting with the enlarged sickle claw on each second toe. Estimated at in length with a body mass about or more, ''Achillobator'' is considered to be one of the largest dromaeosaurs, along with ''Austroraptor'', ''Dakotaraptor'' and ''Utahraptor''. ''Achillobator'' was a ...
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Itemirus
''Itemirus'' is a genus of dromaeosaurid theropod dinosaur from the Turonian age of the Late Cretaceous period of Uzbekistan. Discovery ''Itemirus'' is known from a single small damaged fossil braincase or neurocranium, in 1958 found near the village of Itemir at the Dzharakuduk escarpment in layers of the Bissekty Formation. This holotype has accession number PIN 327/699. The type species, ''Itemirus medullaris'', was named and described by Sergei Kurzanov in 1976. The generic name refers to Itemir. The specific name refers to the medulla oblongata, the brain part encased by the partial braincase.Kurzanov, S. M. (1976) Braincase structure in the carnosaur ''Itemirus'' n. gen. and some aspects of the cranial anatomy of dinosaurs. Paleontological Journal 10:361-369. Classification Kurzanov noted anatomical similarities to the Tyrannosauridae and the Dromaeosauridae; he assigned ''Itemirus'' to a separate Itemiridae. In 2004 Thomas Holtz suggested it was a member of the Tyran ...
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Velociraptor
''Velociraptor'' (; ) is a genus of small dromaeosaurid dinosaur that lived in Asia during the Late Cretaceous epoch, about 75 million to 71 million years ago. Two species are currently recognized, although others have been assigned in the past. The type species is ''V. mongoliensis''; fossils of this species have been discovered in the Djadochta Formation, Mongolia. A second species, ''V. osmolskae'', was named in 2008 for skull material from the Bayan Mandahu Formation, China. Smaller than other dromaeosaurids like ''Deinonychus'' and ''Achillobator'', ''Velociraptor'' was about long with a body mass between . It nevertheless shared many of the same anatomical features. It was a bipedal, feathered carnivore with a long tail and an enlarged sickle-shaped claw on each hindfoot, which is thought to have been used to tackle and restrain prey. ''Velociraptor'' can be distinguished from other dromaeosaurids by its long and low skull, with an upturned snout. ''Velociraptor'' (com ...
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Dakotaraptor
''Dakotaraptor'' (meaning “thief from Dakota”) is a potentially chimaeric genus of large dromaeosaurid theropod dinosaur that lived in North America during the Late Cretaceous period. The remains have been found in the Maastrichtian stage of the Hell Creek Formation, dated to the very end of the Mesozoic era, making ''Dakotaraptor'' one of the last surviving dromaeosaurids. The remains of ''D. steini'' were discovered in a multi-species bonebed. Elements of the holotype and referred specimens were later found to belong to trionychid turtles, and further analysis of potential non-dromaeosaurid affinities of the holotype and referred material have not yet been conducted. Phylogenetic analyses of ''D. steini'' place it in a variety of positions in the Dromaeosauridae. Discovery and naming In 2005, paleontologist Robert DePalma in Harding County, South Dakota discovered a fluvial bonebed bearing the remains of a variety of dinosaurian and non-dinosaurian remains, which yielde ...
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Yurgovuchia
''Yurgovuchia'' (meaning "coyote") is a genus of dromaeosaurid theropod dinosaurs that lived in North America during the Early Cretaceous period in what is now the Cedar Mountain Formation. It contains a single species, ''Yurgovuchia doellingi''. The remains were discovered in Utah, United States. Discovery and naming The holotype was collected by Donald D. DeBlieux in 2005, from Don's Place, part of the Doelling's Bowl bone bed in Grand County, Utah. This bone bed is in the lower Yellow Cat Member of the Cedar Mountain Formation, dating probably to the Valanginian stage of the Early Cretaceous period, about 139–134.6 million years ago. ''Yurgovuchia'' was first described and named by Phil Senter, James I. Kirkland, Donald D. DeBlieux, Scott Madsen and Natalie Toth in 2012 and the type species is ''Yurgovuchia doellingi''. The generic name is derived from the Ute word ''yurgovuch'', meaning coyote, a predator of similar size to ''Y. doellingi'' which currently inhabi ...
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Dromaeosauroides
''Dromaeosauroides'' is a genus of dromaeosaurid theropod dinosaur from the Early Cretaceous of what is now Denmark and possibly also England. It was discovered in the Jydegaard Formation in the Robbedale valley, on the island of Bornholm in the Baltic Sea. This is the only likely place for dinosaur remains to be discovered on Danish territory, since the Mesozoic deposits exposed in the rest of the country are marine. ''Dromaeosauroides'' is the first known dinosaur from this location, and the only one which has been scientifically named. It is one of the oldest known dromaeosaurs in the world, and the first known uncontested dromaeosaur from the Early Cretaceous of Europe. It is known from two teeth, the first of which was found in 2000 and the second in 2008. Based on the first tooth (the holotype), the genus and species ''Dromaeosauroides bornholmensis'' was named in 2003. The genus name means "''Dromaeosaurus''-like", due to the similarity to the teeth of that genus, and t ...
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