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Achillobator
''Achillobator'' ( ; meaning "Achilles hero") is a genus of large dromaeosaurid dinosaurs that lived in Asia during the Late Cretaceous period about 96 million to 89 million years ago in what is now the Bayan Shireh Formation. The genus is currently monotypic, only including the type species ''A. giganticus''. The first remains were found in 1989 during a Mongolian-Russian field expedition in Mongolia and later described in 1999. Remains at the type locality of ''Achillobator'' may represent additional specimens. It represents the first and largest dromaeosaurid known from the Bayan Shireh Formation. It was a large, heavy-built, ground-dwelling, bipedal carnivore that would have been an active feathered predator hunting with the enlarged sickle claw on each second toe. Estimated at in length with a body mass about or more, ''Achillobator'' is considered to be one of the largest dromaeosaurs, along with ''Austroraptor'', ''Dakotaraptor'' and ''Utahraptor''. ''Achillobator'' was a ...
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Dromaeosaurid
Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning 'runner', and ('), meaning 'lizard'. In informal usage, they are often called raptors (after ''Velociraptor''), a term popularized by the film '' Jurassic Park''; a few types include the term "raptor" directly in their name and have come to emphasize their bird-like appearance and speculated bird-like behavior. Dromaeosaurid fossils have been found across the globe in North America, Europe, Africa, Asia, South America and Antarctica, with some fossils giving credence to the possibility that they inhabited Australia as well. They first appeared in the mid-Jurassic Period (late Bathonian stage, about 167 million years ago) and survived until the end of the Cretaceous (Maastrichtian stage, 66 ma), existing until the Cretaceous–Paleogene ex ...
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1999 In Paleontology
Flora Ferns and fern allies Cycads Angiosperms Arthropods Insects Mollusca Newly named bivalves Fish Newly named actinopterygii ("ray-finned fish") Archosauromorphs * Ornithomimid gastroliths documented.Kobayashi et al. (1999). Sanders, Manley, and Carpenter (2001), "Table 12.1" page 167. Newly named dinosauriforms Newly named dinosaurs Data courtesy of George Olshevsky's dinosaur genera list. Newly named birds Newly named pterosaurs Lepidosauromorphs Newly named plesiosaurs Newly named scincomorphans References {{commons category, 1999 in paleontology 1990s in paleontology Paleontology Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
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Dromaeosaurus
''Dromaeosaurus'' (, "running lizard") is a genus of dromaeosaurid theropod dinosaur which lived during the Late Cretaceous period (middle late Campanian and Maastrichtian), sometime between 80 and 69 million years ago, in Alberta, Canada and the western United States. The type species is ''Dromaeosaurus albertensis'', which was described by William Diller Matthew and Barnum Brown in 1922. Its fossils were unearthed in the Dinosaur Park Formation. Teeth attributed to this genus have been found in the Prince Creek Formation. ''Dromaeosaurus'' is the type genus of both Dromaeosauridae and Dromaeosaurinae, which include many genera with similar characteristics to ''Dromaeosaurus'' such as possibly its closest relative '' Dakotaraptor''. ''Dromaeosaurus'' was heavily built, more so than other dromaeosaurs that are similar in size, like ''Velociraptor''. Discovery and naming Despite receiving widespread attention in popular books on dinosaurs, and the usage of a complete mounted skel ...
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Bayan Shireh Formation
The Bayan Shireh Formation (also known as Baynshiree/Baynshire, Baynshirenskaya Svita or Baysheen Shireh) is a geological formation in Mongolia, that dates to the Cretaceous period. It was first described and established by Vasiliev et al. 1959. Description The Bayan Shireh Formation is primarily composed by varicoloured claystones and sandstones with calcareous concretions and characterized by grey mudstones and yellowish-brown medium grained sandstones. Up to thick, the most complete sections are found in the eastern Gobi Desert, consisting of fine-grained, often cross-stratified gray sandstone interbedded with claystone and concretionary, intraformational conglomerates with relatively thick units of red to brown mudstone in the upper part. The Baynshire and Burkhant localities are mainly composed by mudstone, siltstone, sandstone, and conglomerates, with most of their sedimentation being fluvial. The environments that were present on the Bayan Shireh Formation consisted ma ...
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Dakotaraptor
''Dakotaraptor'' (meaning “thief from Dakota”) is a potentially chimaeric genus of large dromaeosaurid theropod dinosaur that lived in North America during the Late Cretaceous period. The remains have been found in the Maastrichtian stage of the Hell Creek Formation, dated to the very end of the Mesozoic era, making ''Dakotaraptor'' one of the last surviving dromaeosaurids. The remains of ''D. steini'' were discovered in a multi-species bonebed. Elements of the holotype and referred specimens were later found to belong to trionychid turtles, and further analysis of potential non-dromaeosaurid affinities of the holotype and referred material have not yet been conducted. Phylogenetic analyses of ''D. steini'' place it in a variety of positions in the Dromaeosauridae. Discovery and naming In 2005, paleontologist Robert DePalma in Harding County, South Dakota discovered a fluvial bonebed bearing the remains of a variety of dinosaurian and non-dinosaurian remains, which yielde ...
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Utahraptor
''Utahraptor'' (meaning "Utah's thief") is a genus of large dromaeosaurid dinosaur that lived in North America during the Early Cretaceous period. It was a heavy-built, ground-dwelling, bipedal carnivore. It contains a single species, ''Utahraptor ostrommaysi'', which is one of the largest-known members of the family Dromaeosauridae, measuring long and weighing . Discovery and naming The first specimens of ''Utahraptor'' were found in 1975 by Jim Jensen in the Dalton Wells Quarry in east-central Utah, near the town of Moab, but did not receive much attention. After a find of a large foot-claw by Carl Limone in October 1991, James Kirkland, Robert Gaston, and Donald Burge uncovered further remains of ''Utahraptor'' in 1991 in the Gaston Quarry in Grand County, Utah, within the Yellow Cat and Poison Strip members of the Cedar Mountain Formation. The holotype of ''Utahraptor'', CEUM 184v.86 consists of a second pedal ungual, with potentially assigned elements from other specim ...
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Altangerel Perle
Altangerel Perle (born 1945) is a Mongolian palaeontologist. He is employed at the National University of Mongolia. He has described species such as ''Goyocephale lattimorei'', '' Achillobator giganticus'' and '' Erlikosaurus andrewsi''. He has been honored by Polish palaeontologist Halszka Osmólska Halszka Osmólska (September 15, 1930 – March 31, 2008) was a Polish paleontologist who had specialized in Mongolian dinosaurs. Biography She was born in 1930 in Poznań. In 1949, she began to study biology at Faculty of Biology and Earth Scie ..., who named the species '' Hulsanpes perlei'' after him. References 1945 births Living people Mongolian paleontologists National University of Mongolia faculty {{Mongolia-bio-stub ...
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Eudromaeosauria
Eudromaeosauria ("true dromaeosaurs") is a subgroup of terrestrial dromaeosaurid theropod dinosaurs. They were relatively small to medium-sized, feathered hypercarnivores (with diets consisting almost entirely of other terrestrial vertebrates) that flourished in the Cretaceous Period. Eudromaeosaur fossils are known almost exclusively from the northern hemisphere. They first appeared in the early Cretaceous Period (early Aptian stage, about 124 million years ago) and survived until the end of the Cretaceous (Maastrichtian stage, Ma). The earliest known definitive eudromaeosaur is the dromaeosaurine '' Utahraptor ostrommaysi'', from the Cedar Mountain Formation, dated to 124 million years ago. However, the earlier (143-million-year-old) fossils such as those of '' Nuthetes destructor'' and several indeterminate teeth dating to the Kimmeridgian stage may represent eudromaeosaurs. Description While other dromaeosaurids filled a variety of specialized ecological niches, mainly tho ...
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Austroraptor
''Austroraptor'' ( ) is a genus of large dromaeosaurid dinosaur that lived during the Late Cretaceous Period in what is now Argentina. ''Austroraptor'' was a large-sized, moderately-built, ground-dwelling, bipedal carnivore, estimated at in length. It is one of the largest dromaeosaurids known, with only ''Achillobator'', ''Dakotaraptor'', and ''Utahraptor'' approaching or surpassing it in length. Discovery and naming The type specimen of ''Austroraptor cabazai'', holotype MML-195, was recovered in the Bajo de Santa Rosa locality of the Allen Formation, in Río Negro, Argentina. The specimen was collected in 2002 by the team of Fernando Emilio Novas of the '' Museo Argentino de Ciencias Naturales''. It consists of a fragmentary skeleton including parts of the skull, lower jaw, a few neck and torso vertebrae, some ribs, a humerus, and assorted bones from both legs. The specimen was prepared by Marcelo Pablo Isasi and Santiago Reuil. In 2008, the type species ''Austroraptor cab ...
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Sediment
Sediment is a naturally occurring material that is broken down by processes of weathering and erosion, and is subsequently transported by the action of wind, water, or ice or by the force of gravity acting on the particles. For example, sand and silt can be carried in suspension in river water and on reaching the sea bed deposited by sedimentation; if buried, they may eventually become sandstone and siltstone (sedimentary rocks) through lithification. Sediments are most often transported by water (fluvial processes), but also wind (aeolian processes) and glaciers. Beach sands and river channel deposits are examples of fluvial transport and deposition, though sediment also often settles out of slow-moving or standing water in lakes and oceans. Desert sand dunes and loess are examples of aeolian transport and deposition. Glacial moraine deposits and till are ice-transported sediments. Classification Sediment can be classified based on its grain size, grain shape, and c ...
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Maxilla
The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxillary bones are fused at the intermaxillary suture, forming the anterior nasal spine. This is similar to the mandible (lower jaw), which is also a fusion of two mandibular bones at the mandibular symphysis. The mandible is the movable part of the jaw. Structure In humans, the maxilla consists of: * The body of the maxilla * Four processes ** the zygomatic process ** the frontal process of maxilla ** the alveolar process ** the palatine process * three surfaces – anterior, posterior, medial * the Infraorbital foramen * the maxillary sinus * the incisive foramen Articulations Each maxilla articulates with nine bones: * two of the cranium: the frontal and ethmoid * seven of the face: the nasal, zygomatic, lacrimal, inferior n ...
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Sandstone
Sandstone is a clastic sedimentary rock composed mainly of sand-sized (0.0625 to 2 mm) silicate grains. Sandstones comprise about 20–25% of all sedimentary rocks. Most sandstone is composed of quartz or feldspar (both silicates) because they are the most resistant minerals to weathering processes at the Earth's surface. Like uncemented sand, sandstone may be any color due to impurities within the minerals, but the most common colors are tan, brown, yellow, red, grey, pink, white, and black. Since sandstone beds often form highly visible cliffs and other topographic features, certain colors of sandstone have been strongly identified with certain regions. Rock formations that are primarily composed of sandstone usually allow the percolation of water and other fluids and are porous enough to store large quantities, making them valuable aquifers and petroleum reservoirs. Quartz-bearing sandstone can be changed into quartzite through metamorphism, usually related to ...
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