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Dorsal Trigeminothalamic Tract
The dorsal trigeminal tract, dorsal trigeminothalamic tract, or posterior trigeminothalamic tract, is composed of second-order neuronal axons. These fibers carry sensory information about discriminative touch and conscious proprioception in the oral cavity from the principal (chief sensory) nucleus of the trigeminal nerve to the ventral posteromedial (VPM) nucleus of the thalamus. The dorsal trigeminothalamic tract is also called the posterior trigeminal lemniscus. Pathway The first-order neurons (from the trigeminal ganglion) enter the pons and synapse on second-order neurons in the principal (chief sensory) nucleus. Axons of the second-order neurons then decussate to enter the trigeminal lemniscus in the midbrain and then ascend to the ventral posteromedial nucleus of the contralateral thalamus, forming the ventral trigeminothalamic tract. A subset of these fibers do not decussate and travel to the ipsilateral ventral posteromedial nucleus of the thalamus. These non-decu ...
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Second-order Neuron
In physiology, the somatosensory system is the network of neural structures in the brain and body that produce the perception of touch ( haptic perception), as well as temperature ( thermoception), body position (proprioception), and pain. It is a subset of the sensory nervous system, which also represents visual, auditory, olfactory, and gustatory stimuli. Somatosensation begins when mechano- and thermosensitive structures in the skin or internal organs sense physical stimuli such as pressure on the skin (see mechanotransduction, nociception). Activation of these structures, or receptors, leads to activation of peripheral sensory neurons that convey signals to the spinal cord as patterns of action potentials. Sensory information is then processed locally in the spinal cord to drive reflexes, and is also conveyed to the brain for conscious perception of touch and proprioception. Note, somatosensory information from the face and head enters the brain through peripheral s ...
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Proprioception
Proprioception ( ), also referred to as kinaesthesia (or kinesthesia), is the sense of self-movement, force, and body position. It is sometimes described as the "sixth sense". Proprioception is mediated by proprioceptors, mechanosensory neurons located within muscles, tendons, and joints. Most animals possess multiple subtypes of proprioceptors, which detect distinct kinematic parameters, such as joint position, movement, and load. Although all mobile animals possess proprioceptors, the structure of the sensory organs can vary across species. Proprioceptive signals are transmitted to the central nervous system, where they are integrated with information from other sensory systems, such as the visual system and the vestibular system, to create an overall representation of body position, movement, and acceleration. In many animals, sensory feedback from proprioceptors is essential for stabilizing body posture and coordinating body movement. System overview In vertebrates, limb ve ...
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Oral Cavity
In animal anatomy, the mouth, also known as the oral cavity, or in Latin cavum oris, is the opening through which many animals take in food and issue vocal sounds. It is also the cavity lying at the upper end of the alimentary canal, bounded on the outside by the lips and inside by the pharynx. In tetrapods, it contains the tongue and, except for some like birds, teeth. This cavity is also known as the buccal cavity, from the Latin ''bucca'' ("cheek"). Some animal phyla, including arthropods, molluscs and chordates, have a complete digestive system, with a mouth at one end and an anus at the other. Which end forms first in ontogeny is a criterion used to classify bilaterian animals into protostomes and deuterostomes. Development In the first multicellular animals, there was probably no mouth or gut and food particles were engulfed by the cells on the exterior surface by a process known as endocytosis. The particles became enclosed in vacuoles into which enzymes were secre ...
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Principal Sensory Nucleus Of Trigeminal Nerve
The principal sensory nucleus of trigeminal nerve (or chief sensory nucleus of V, main trigeminal sensory nucleus) is a group of second-order neurons which have cell bodies in the caudal pons. It receives information about discriminative sensation and light touch of the face as well as conscious proprioception of the jaw via first order neurons of CN V. * Most of the sensory information crosses the midline and travels to the contralateral ventral posteromedial nucleus (VPM) of the thalamus via the anterior trigeminothalamic tract. * However, information of the ''oral cavity'' travels to the ipsilateral VPM of the thalamus via the dorsal trigeminothalamic tract The dorsal trigeminal tract, dorsal trigeminothalamic tract, or posterior trigeminothalamic tract, is composed of second-order neuronal axons. These fibers carry sensory information about discriminative touch and conscious proprioception in the o .... {{Authority control Cranial nerve nuclei Trigeminal nerve Pons ...
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Ventral Posteromedial Nucleus
The ventral posteromedial nucleus (VPM) is a nucleus of the thalamus. Inputs and outputs The VPM contains synapses between second and third order neurons from the anterior (ventral) trigeminothalamic tract and posterior (dorsal) trigeminothalamic tract. These neurons convey sensory information from the face and oral cavity. Third order neurons in the trigeminothalamic systems project to the postcentral gyrus. The VPM also receives taste afferent information from the solitary tract The solitary tract (tractus solitarius, or fasciculus solitarius), is a compact fiber bundle that extends longitudinally through the posterolateral region of the medulla oblongata. The solitary tract is surrounded by the solitary nucleus, and des ... and projects to the cortical gustatory area. Subareas VPMpc The parvicellular part of the ventroposterior medial nucleus (VPMpc) is argued by some as not an actually part of the VPM, because it does not project to the somatosensory cortex as the r ...
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Thalamus
The thalamus (from Greek θάλαμος, "chamber") is a large mass of gray matter located in the dorsal part of the diencephalon (a division of the forebrain). Nerve fibers project out of the thalamus to the cerebral cortex in all directions, allowing hub-like exchanges of information. It has several functions, such as the relaying of sensory signals, including motor signals to the cerebral cortex and the regulation of consciousness, sleep, and alertness. Anatomically, it is a paramedian symmetrical structure of two halves (left and right), within the vertebrate brain, situated between the cerebral cortex and the midbrain. It forms during embryonic development as the main product of the diencephalon, as first recognized by the Swiss embryologist and anatomist Wilhelm His Sr. in 1893. Anatomy The thalamus is a paired structure of gray matter located in the forebrain which is superior to the midbrain, near the center of the brain, with nerve fibers projecting out to the ...
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Trigeminal Lemniscus
The trigeminal lemniscus, also called the trigeminothalamic tract, is composed of the ventral trigeminal tract, and the dorsal trigeminal tract – nerve tracts that convey tactile, pain, and temperature impulses from the skin of the face, the mucous membranes of the nasal and oral cavities, and the eye, as well as proprioceptive information from the facial and masticatory muscles. The trigeminal lemniscus is composed of second order neuronal axons in the brainstem. It carries sensory information from the trigeminal system to the ventral posteromedial (VPM) nucleus of the thalamus. This tract was historically considered a cephalic division of the medial lemniscus due to the close proximity of the two ascending tracts. Like the medial lemniscus in the dorsal column-medial lemniscus pathway (DCML), that carries mechanosensory information from part of the head and the rest of the body, the trigeminal lemniscus carries mechanosensory information from the face. However, the trigemin ...
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Dorsal Root Ganglion
A dorsal root ganglion (or spinal ganglion; also known as a posterior root ganglion) is a cluster of neurons (a ganglion) in a dorsal root of a spinal nerve. The cell bodies of sensory neurons known as first-order neurons are located in the dorsal root ganglia. The axons of dorsal root ganglion neurons are known as afferents. In the peripheral nervous system, afferents refer to the axons that relay sensory information into the central nervous system (i.e. the brain and the spinal cord). Structure The neurons comprising the dorsal root ganglion are of the pseudo-unipolar type, meaning they have a cell body (soma) with two branches that act as a single axon, often referred to as a ''distal process'' and a ''proximal process''. Unlike the majority of neurons found in the central nervous system, an action potential in posterior root ganglion neuron may initiate in the ''distal process'' in the periphery, bypass the cell body, and continue to propagate along the ''proximal process ...
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Midbrain
The midbrain or mesencephalon is the forward-most portion of the brainstem and is associated with vision, hearing, motor control, sleep and wakefulness, arousal (alertness), and temperature regulation. The name comes from the Greek ''mesos'', "middle", and ''enkephalos'', "brain". Structure The principal regions of the midbrain are the tectum, the cerebral aqueduct, tegmentum, and the cerebral peduncles. Rostrally the midbrain adjoins the diencephalon (thalamus, hypothalamus, etc.), while caudally it adjoins the hindbrain (pons, medulla and cerebellum). In the rostral direction, the midbrain noticeably splays laterally. Sectioning of the midbrain is usually performed axially, at one of two levels – that of the superior colliculi, or that of the inferior colliculi. One common technique for remembering the structures of the midbrain involves visualizing these cross-sections (especially at the level of the superior colliculi) as the upside-down face of a be ...
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Ventral Posteromedial Nucleus
The ventral posteromedial nucleus (VPM) is a nucleus of the thalamus. Inputs and outputs The VPM contains synapses between second and third order neurons from the anterior (ventral) trigeminothalamic tract and posterior (dorsal) trigeminothalamic tract. These neurons convey sensory information from the face and oral cavity. Third order neurons in the trigeminothalamic systems project to the postcentral gyrus. The VPM also receives taste afferent information from the solitary tract The solitary tract (tractus solitarius, or fasciculus solitarius), is a compact fiber bundle that extends longitudinally through the posterolateral region of the medulla oblongata. The solitary tract is surrounded by the solitary nucleus, and des ... and projects to the cortical gustatory area. Subareas VPMpc The parvicellular part of the ventroposterior medial nucleus (VPMpc) is argued by some as not an actually part of the VPM, because it does not project to the somatosensory cortex as the r ...
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Anterior Trigeminothalamic Tract
The ventral trigeminal tract, ventral trigeminothalamic tract, anterior trigeminal tract, or anterior trigeminothalamic tract, is a tract composed of second order neuronal axons. These fibers carry sensory information about discriminative and crude touch, conscious proprioception, pain, and temperature from the head, face, and oral cavity. The ventral trigeminal tract connects the two major components of the brainstem trigeminal complex – the principal, or main sensory nucleus and the spinal trigeminal nucleus, to the ventral posteromedial nucleus of the thalamus. The ventral trigeminal tract is also called the anterior trigeminal lemniscus. Structure The first order neurons (from the trigeminal ganglion) enter the pons and synapse in the principal (chief sensory) nucleus or spinal trigeminal nucleus. Axons of the second order neurons cross the midline and terminate in the ventral posteromedial nucleus of the contralateral thalamus (as opposed to the ventral posterolateral nucl ...
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Sensory Cortex
The sensory cortex can refer informally to the primary somatosensory cortex, or it can be used as a term for the primary and secondary cortices of the different senses (two cortices each, on left and right hemisphere): the visual cortex on the occipital lobes, the auditory cortex on the temporal lobes, the primary olfactory cortex on the uncus of the piriform region of the temporal lobes, the gustatory cortex on the insular lobe (also referred to as the insular cortex), and the primary somatosensory cortex on the anterior parietal lobes. Just posterior to the primary somatosensory cortex lies the somatosensory association cortex, which integrates sensory information from the primary somatosensory cortex (temperature, pressure, etc.) to construct an understanding of the object being felt. Inferior to the frontal lobes are found the olfactory bulbs, which receive sensory input from the olfactory nerves and route those signals throughout the brain. Not all olfactory information is rou ...
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