Domeykosaurus
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Domeykosaurus
''Arackar'' (meaning "skeleton" in Kunza) is an extinct genus of lithostrotian sauropod, possibly part of the Saltasauridae, discovered in the Hornitos Formation of Atacama Province, Chile. The genus contains a single species, ''Arackar licanantay'', described by Rubilar-Rogers ''et al.'' in 2021. Discovery and naming The holotype was discovered in 1993 at Quebrada La Higuera, approximately 75 km (47 mi) south of Copiapó. The fossil material was briefly mentioned by Rubilar-Rogers & Gutstein in 2012 and was found to not be referable to the genus ''Atacamatitan''. In 2021, ''Arackar licanantay'' was named and described by David Rubilar-Rogers, Alexander O. Vargas, Bernardo González Riga, Sergio Soto-Acuña, Jhonatan Alarcón-Muñoz, José Iriarte-Díaz, Carlos Arévalo and Carolina S. Gutstein. The holotype, SNGM-1, was found in a layer of the Hornitos Formation in Chile that dates back to the Late Cretaceous period ( Campanian-Maastrichtian stages). It includes vertebrae of t ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Ant ...
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Femur
The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with the tibia (shinbone) and patella (kneecap), forming the knee joint. By most measures the two (left and right) femurs are the strongest bones of the body, and in humans, the largest and thickest. Structure The femur is the only bone in the upper leg. The two femurs converge medially toward the knees, where they articulate with the proximal ends of the tibiae. The angle of convergence of the femora is a major factor in determining the femoral-tibial angle. Human females have thicker pelvic bones, causing their femora to converge more than in males. In the condition ''genu valgum'' (knock knee) the femurs converge so much that the knees touch one another. The opposite extreme is ''genu varum'' (bow-leggedness). In the general populatio ...
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Baurutitan
''Baurutitan'' is a genus of sauropod dinosaur that lived during the Late Cretaceous in what is now Brazil. The type species, ''Baurutitan britoi'', was described in 2005 by Kellner and colleagues, although the fossil remains had already been discovered in 1957. ''Baurutitan'' is classified as a lithostrotian titanosaur, and is distinguished from related genera based on its distinctive caudal vertebrae. This South American dinosaur was found in the Serra da Galga Formation near Uberaba, in the Brazilian state of Minas Gerais. Discovery The holotype of ''Baurutitan'' were found in 1957 by Llewellyn Ivor Price, the famous Brazilian paleontologist, in the region of Peirópolis, Minas Gerais. However, it was not until 2005 that ''Baurutitan'' was officially published and named. The works of Price in Peirópolis began in 1947 after Jesuíno Felicíssimo Junior, from the Instituto Geográfico e Geológico of São Paulo, told him about the presence of fossils in the region. Price then ...
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Malawisaurus
''Malawisaurus'' (meaning "Malawi lizard") is an extinct genus of titanosaurian sauropod dinosaur. It is known from the Dinosaur Beds of northern Malawi, which probably date to the Aptian stage of the Early Cretaceous. The type species is ''M. dixeyi'' and the specific name honours Frederick Augustus Dixey. Discovery and naming ''Malawisaurus dixeyi'' was originally described in 1928 by Sidney H. Haughton as a species of ''Gigantosaurus'' (an invalid name for the diplodocid currently known as ''Tornieria''). Haughton considered it closely related to the species ''G. robustus'' (later the type species of ''Janenschia''). The holotype was discovered in the "Dinosaur Beds" of Malawi (then known as the Nyasaland Protectorate), which are usually considered to be of Barremian-Aptian age based on K–Ar dating, though they have also been suggested to be Late Cretaceous in age based on the vertebrate assemblage, and possibly also the Lupata Group. In 1993 it was placed in the newly na ...
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Rapetosaurus
''Rapetosaurus'' ( ) is a genus of titanosaurian sauropod dinosaur that lived in Madagascar from 70 to 66 million years ago, at the end of the Cretaceous Period. Only one species, ''Rapetosaurus krausei'', has been identified. Like other sauropods, ''Rapetosaurus'' was a quadrupedal herbivore; it is calculated to have reached lengths of 15 metres (49 ft). Description ''Rapetosaurus'' was a fairly typical sauropod, with a short and slender tail, a very long neck and a huge, elephant-like body. Its head resembles the head of a diplodocid, with a long, narrow snout and nostrils on the top of its skull. It was a herbivore and its small, pencil-like teeth were good for ripping the leaves off trees but not for chewing. It was fairly modest in size, for a titanosaur. The juvenile specimen measured from head to tail, and "probably weighed about as much as an elephant". An adult would have been about twice as long ( in length) which is still less than half the length of its ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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Isisaurus
''Isisaurus'' (named after the Indian Statistical Institute) is a genus of titanosaurian dinosaur from the Late Cretaceous Lameta Formation of India. The genus contains a single species, ''Isisaurus colberti''. Discovery and Naming The type specimen of ''Isisaurus colberti'', ISI R 335/1-65, was originally described and named as ''Titanosaurus colberti'' by Sohan Lal Jain and Saswati Bandyopadhyay in 1997. The specific name honours Edwin Harris Colbert. In 2003, the fossils were designated as belonging to its own genus by Wilson and Upchurch. The generic name, "''Isisaurus''," combines a reference to the Indian Statistical Institute (ISI) with the Greek "saurus," meaning "lizard." It had a short, vertically directed neck and long forelimbs, making it considerably different from other sauropods. The humerus is 148 centimetres long. ''Isisaurus'' is known from better remains than many other titanosaurs that were known at the time of its description. Much of its postcranial ske ...
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Sister Taxon
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxonomi ...
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Lithostrotia
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of ''Malawisaurus'' and ''Saltasaurus'' and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek , meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch ''et al.'' include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous. History of research In 1895, Richard Lydekker named the family Titanosauridae to summarize sauropods with procoelous (concave on the front) caudal vertebrae. The name Titanosauridae has since been widely used, and was defined by Salgado and colleagues (1997), Gonzalaz-Riga (2003), and Salgado (2003) as a node-based ...
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Titanosauria
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as ''Patagotitan''—estimated at long with a weight of —and the comparably-sized ''Argentinosaurus'' and ''Puertasaurus'' from the same region. The group's name alludes to the mythological Titans of ancient Greek mythology, via the type genus (now considered a ''nomen dubium)'' ''Titanosaurus''. Together with the brachiosaurids and relatives, titanosaurs make up the larger sauropod clade Titanosauriformes. Titanosaurs have long been a poorly-known group, and the relationships between titanosaur species are still not well-understood. Description Titanosauria have the largest ran ...
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Neural Spine
The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic irregular bone whose complex structure is composed primarily of bone, and secondarily of hyaline cartilage. They show variation in the proportion contributed by these two tissue types; such variations correlate on one hand with the cerebral/caudal rank (i.e., location within the backbone), and on the other with phylogenetic differences among the vertebrate taxa. The basic configuration of a vertebra varies, but the bone is its ''body'', with the central part of the body constituting the ''centrum''. The upper (closer to) and lower (further from), respectively, the cranium and its central nervous system surfaces of the vertebra body support attachment to the intervertebral discs. The posterior part of a vertebra forms a vertebral arch (in ...
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Arackar Vetrebrae
''Arackar'' (meaning "skeleton" in Kunza) is an extinct genus of lithostrotian sauropod, possibly part of the Saltasauridae, discovered in the Hornitos Formation of Atacama Province, Chile. The genus contains a single species, ''Arackar licanantay'', described by Rubilar-Rogers ''et al.'' in 2021. Discovery and naming The holotype was discovered in 1993 at Quebrada La Higuera, approximately 75 km (47 mi) south of Copiapó. The fossil material was briefly mentioned by Rubilar-Rogers & Gutstein in 2012 and was found to not be referable to the genus ''Atacamatitan''. In 2021, ''Arackar licanantay'' was named and described by David Rubilar-Rogers, Alexander O. Vargas, Bernardo González Riga, Sergio Soto-Acuña, Jhonatan Alarcón-Muñoz, José Iriarte-Díaz, Carlos Arévalo and Carolina S. Gutstein. The holotype, SNGM-1, was found in a layer of the Hornitos Formation in Chile that dates back to the Late Cretaceous period ( Campanian-Maastrichtian stages). It includes vertebrae of ...
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