Discocerid
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Discocerid
Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck. Evolution The origin of the Discosorida is unknown, thought at one timeRousseau H. Flower. 1964. The Nautiloid Order Ellesmerocerida (Cephalopods); relevant pages. Memoir 12, ...
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Cephalopod
A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by ''Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been ident ...
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Cyrtogomphoceratidae
The Cyrtogomphoceratidae are a family in the cephalopod order Discosorida that comprises genera commonly with compressed, endogastrically curved shells. Siphuncles lie close to the ventral side, segments are broadly inflated, connecting rings thick and apically expanded thick bullettes. Chambers are short, separated by shallow, dish shaped septa. Apertures are generally simple. The Cyrtogomphoceratidae are derived from the discosorid family Reudemannoceratidae, probably from '' Reudemannoceras'', through the ancestral genus '' Ulrichoceras'', and have a range from the Middle Ordovician to the Lower Silurian. The familyTeicher, C. 1964. Nautiloidea-Discosorida. Treatise on Invertebrate Paleontology, Part K. Teichert and Moore, eds. includes: :*'' Cyrtogomphoceras'' :*'' Kiaeroceras'' :*'' Konglungenoceras'' :*'' Landeroceras'' :*'' Parryoceras'' :*'' Strandoceras'' :*'' Ulrichoceras'' ''Ulrichoceras'' is also considered the source for the exogastric Westonoceratidae. T ...
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Cephalopod Orders
A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by ''Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been identified. Tw ...
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Discosorida
Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck. Evolution The origin of the Discosorida is unknown, thought at one timeRousseau H. Flower. 1964. The Nautiloid Order Ellesmerocerida (Cephalopods); relevant pages. Memoir 12, Ne ...
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Oncocerida
The Oncocerida comprise a diverse group of generally small nautiloid cephalopods known from the Middle Ordovician to the Mississippian (early Carboniferous; one possible member is known from the Early Permian), in which the connecting rings are thin and siphuncle segments are variably expanded (Flower, 1950). At present the order consists of some 16 families, a few of which, such as the Oncoceratidae, Brevicoceratidae, and Acleistoceratidae contain a fair number of genera each while others like the Trimeroceratidae and Archiacoceratidae are represented by only two or three (Sweet, 1964). Physical characteristics The shells of oncocerids are primarily somewhat compressed cyrtoconic brevicones. More advanced forms include gyrocones, serpenticones, torticones, and elongate orthocones and cyrtocones, reflective of the different families and genera (Flower, 1950; Sweet, 1964). The siphuncle in the Oncocerida is commonly located at or near the ventral margin. Connecting rings are mo ...
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Benthic
The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "the depths." Organisms living in this zone are called benthos and include microorganisms (e.g., bacteria and fungi) as well as larger invertebrates, such as crustaceans and polychaetes. Organisms here generally live in close relationship with the substrate and many are permanently attached to the bottom. The benthic boundary layer, which includes the bottom layer of water and the uppermost layer of sediment directly influenced by the overlying water, is an integral part of the benthic zone, as it greatly influences the biological activity that takes place there. Examples of contact soil layers include sand bottoms, rocky outcrops, coral, and bay mud. Description Oceans The benthic region of the ocean begins at the shore line (intertidal ...
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Mandaloceratidae
Mandaloceratidae is a family in the nautiloid cephalopod order Discosorida, from the Middle and Upper(?) Silurian characterized by short, essentially straight shells referred to as breviconic, typically with a faintly exogastric shape produced by the profile of the body chamber. Apertures vary from round to T-shaped, with a long narrow, ventral, hyponomic sinus for the maneuvering funnel. Siphuncles are commonly central or subcentral with generally broad, expanded, segments and usually thin connecting rings. The narrow hyponomic sinus, which is an opening for the water-jet funnel by which the animal could move, much as with modern squid True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fitting t ... and octopods, points toward the lower or ventral side. The cross bar of the "T" or the round ...
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Discosoridae
Discosoridae comprise a family of endogastric discosorids, (Cephalopoda, Nautiloidea), with endocones in the siphuncle, ranging from the Middle Silurian to Middle Devonian. Shell morphology The shell in the Discosoridae is conical, expanding variably with respect to genus. Curvature tends to be slight with the ventral or siphuncular side slightly concave in profile (the endogastric condition), and the opposite dorsal side arched or convex in profile (another endogastric condition). The siphuncle is composed of broadly rounded segments the increase rapidly in size toward the aperture, with growth. Expansion is commonly greater than that of the shell itself, and lies ventral of the center. Septal necks are short, brims long and recumbent. Connecting rings are thin and poorly known. Bullettes at the apical end of the connecting rings, which grasp the periphery of the previous septal foramina and connect to the inside of the previous septal necks, are never swollen. Endocones ar ...
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Lowoceratidae
The Lowoceratidae is a small family of discosorids, early nautiloid cephalopods, from the Middle Silurian in which the characteristic bullette is found only in early growth stages. Lowoceratids were first found in Southampton Island in the Canadian arctic. The Lowoceratidae consists so far of only two genera, ''Lowoceras'' and ''Tuylocras'', named by Foerste and Savage in 1927 and combined by Flower in 1940 as the Lowoceratidae. Both are strongly compressed, exogastric brevicones with the ventral side more narrowly rounded than the dorsal. Shells enlarge moderately to the middle of the mature body chamber, then contract gently toward the aperture. The siphuncle in each is slightly removed from the ventral margin. The siphuncle of ''Tuyloceras'', genotype ''T. percurvatum'', starts off with slender, subquadrate segments with thick rings and swollen bullettes, but later in life the segments become expanded, rings thinner, and bullettes simple and unswollen. ''Lowoceras'', gen ...
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Phragmoceratidae
The Phragmoceratidae is a family of extinct nautiloid cephalopods from the Order Discosorida that lived during the latter part of the Silurian. Diagnosis Phragmoceratids are characterized by generally compressed, upwardly curved endogastric shells with slit-like apertures for egress and ventral siphuncles with broadly expanded segments, thick connecting rings, and small to vestigial bullettes. Phragmoceratids are morphologically similar in regards to their constricted apertures to the more cylindrical and orthoconic Mandaloceratidae, also discosorids, and to the oncocerid Hemiphragmoceratidae Hemiphragmoceratidae is a family of endogastrically brevconic oncocerids characterized by elaborately visored apertures in which the hyponomic sinus in mature specimens is on a spout-like process and there may be lateral and dorsal salients. (S ... (Sweet 1964); both which also come from the Silurian. Derivation The Phragmoceratidae are derived from the Ordovician Cyrtogomphocerat ...
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Westonoceratidae
Westonoceratidae are exogastric, mostly compressed, Discosorida of moderate size from the Middle Ordovician to the Lower Silurian. The siphuncle is typically close to the convexly curved outer margin of the phragmocone – the chambered part of the shell – taken to be ventral but may be more central in some genera. Connecting rings are thin to moderately thick with inflated bullettes grasping the previous septal foremina. Parietal deposits within the siphuncle from endocones in advanced genera and cameral deposits are found in some. The Westonoceratidae are derived from the Cyrtogomphoceratidae through the cyrtogomphoceratid, '' Ulrichoceras''. In turn, the Westonoceratidae gave rise to a small group of discosorids, the Lowoceratidae, which form an intermediary step before the Discosoridae. Two principal lineages have been identified, both stemming from the genus '' Teichertoceras'', a derivative of ''Ulrichoceras''. One goes simply from '' Westonoceras'' to '' Winnipeg ...
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