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Diplognathus
''Diplognathus'' is a genus of arthrodire placoderm from the Late Famennian Cleveland Shale of Late Devonian Ohio, known only from incomplete fragments of jaws and skulls. What fragments are known suggest that the living animals were large-eyed piscivore A piscivore () is a carnivorous animal that eats primarily fish. The name ''piscivore'' is derived . Piscivore is equivalent to the Greek-derived word ichthyophage, both of which mean "fish eater". Fish were the diet of early tetrapod evoluti ...s with weak, but widely gaping jaws. ''D. mirabilis'' is thought to be fairly large, with infragnathals up to 45 centimeters in length. The second species, ''D. larfargei'', was much smaller, with inferognathals averaging about 4 centimeters in length. In 1967, Obruchev placed this genus within Hadrosteidae, on the basis of how the two genera have similar denticle ("teeth") patterns of the inferognathals, though Denison (1978) contested this placement, preferring to leave the ...
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Hadrosteidae
Hadrosteidae is a family of arthrodire placoderms from the Late Devonian. It was originally erected for the late Frasnian '' Hadrosteus'', from the Kellwasserkalk facies. Later, the family was subjectively subsumed into Dinichthyidae due to ''Hadrosteus''' anatomical similarities with '' Dunkleosteus'' and ''Dinichthys''. In 1967, Obruchev placed the enigmatic arthrodire '' incertae sedis'' ''Diplognathus'', from the Eastern American Famennian, within Hadrosteidae, on the basis of how the two genera have similar denticle ("teeth") patterns of the inferognathals, though, Denison (1978) contests this placement. With the redefining of Dinichthyidae as a monotypic family for ''Dinichthys'', and only a few other genera placed within Dunkleosteidae Dunkleosteidae is an extinct family of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator ''Dunkleosteus terrelli'' is the best known member of this group. Phylogeny While members of Dunkleosteida ...
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Arthrodira
Arthrodira (Greek for "jointed neck") is an order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of Placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are protected by a bony ring, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus ''Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed ''Rolfosteus'' measured just ...
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Arthrodira Enigmatic Taxa
Arthrodira (Greek for "jointed neck") is an order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of Placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are protected by a bony ring, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus '' Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, '' Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed '' Rolfosteus'' measured ...
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Cleveland Shale
The Cleveland Shale, also referred to as the Cleveland Member, is a shale geologic formation in the eastern United States. Identification and name The Cleveland Shale was identified in 1870 and named for the city of Cleveland, Ohio. John Strong Newberry, director of the Ohio State Geological Survey, first identified the formation in 1870. He called it the "Cleveland Shale" and designated its type locality at Doan Brook near Cleveland. Details of the type locality and of stratigraphic nomenclature for this unit as used by the U.S. Geological Survey are available on-line at the National Geologic Map Database. Description The primary minerals in the Cleveland Shale are chlorite, illite, pyrite, and quartz. Underground, the Cleveland Shale is black, dull grayish-black, bluish-black, or brownish-black in color. In exposed outcrops, it weathers to red, reddish-brown, or medium brown. Highly weathered rock turns gray. It is fairly fissile, breaking into thin, irregularly shaped ...
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Arthrodire
Arthrodira (Greek for "jointed neck") is an Order (biology), order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of Placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are protected by a bony ring, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus ''Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed ''Rolfosteus ...
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Placodermi
Placodermi (from Greek πλάξ 'plate' and δέρμα 'skin', literally 'Plate (animal anatomy), plate-skinned') is a Class (biology), class of armoured prehistoric fish, known from fossils, which lived from the Silurian to the end of the Devonian period. Their head and thorax were covered by articulated armoured plates and the rest of the body was scale (zoology), scaled or naked, depending on the species. Placoderms were among the first jawed fish; their Fish jaw, jaws likely evolved from the first of their gill arches. Placoderms are thought to be paraphyly, paraphyletic, consisting of several distinct Outgroup (cladistics), outgroups or sister taxon, sister taxa to all living jawed vertebrates, which originated among their ranks. In contrast, one 2016 analysis concluded that placodermi are likely monophyletic, though these analyses have been further dismissed with more transitional taxa between placoderms and modern gnathosthomes, solidifying their paraphyletic status. Plac ...
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Piscivore
A piscivore () is a carnivorous animal that eats primarily fish. The name ''piscivore'' is derived . Piscivore is equivalent to the Greek-derived word ichthyophage, both of which mean "fish eater". Fish were the diet of early tetrapod evolution (via water-bound amphibians during the Devonian period); insectivory came next; then in time, the more terrestrially adapted reptiles and synapsids evolved herbivory. Almost all predatory fishes (most sharks, tuna, billfishes, pikes etc.) are obligated piscivores. Some non-piscine aquatic animals, such as whales, sea lion and crocodilians, are not completely piscivorous, often also preying on invertebrates, marine mammals, waterbirds and even wading land animals in addition to fish; while others, such as the bulldog bat and gharial, are strictly dependent on fish for food. Some creatures, including cnidarians, octopuses, squid, spiders, cetaceans, grizzly bears, jaguars, wolves, snakes, turtles and sea gulls, may have fish as signific ...
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Famennian Life
The Famennian is the latter of two faunal stages in the Late Devonian Epoch. The most recent estimate for its duration estimates that it lasted from around 371.1 million years ago to 359.3 million years ago. An earlier 2012 estimate, still used by the International Commission on Stratigraphy, estimated that it lasted from million years ago to million years ago. It was preceded by the Frasnian stage and followed by the Tournaisian stage. Major events In the seas, a novel major group of ammonoid cephalopods called clymeniids appeared, underwent tremendous diversification and spread worldwide, then just as suddenly went extinct. The beginning of the Famennian is marked by the final stages of a major extinction event, the Kellwasser Event, which is the largest component of the Late Devonian Mass extinction. The end of the Famennian experiences a smaller but still quite severe extinction event, the Hangenberg Event. A brief episode of glaciation, possibly linked to the Hangenber ...
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Paleontology In Ohio
Paleontology in Ohio refers to paleontological research occurring within or conducted by people from the U.S. state of Ohio. Ohio is well known for having a great quantity and diversity of fossils preserved in its Rock (geology), rocks. The state's fossil record begins early in the Paleozoic era, during the Cambrian period. Ohio was generally covered by seawater from that time on through the rest of the early Paleozoic. Local invertebrates included brachiopods, cephalopods, coral, graptolites, and trilobites. Vertebrates included bony fishes and sharks. The first land plants in the state grew during the Devonian. During the Carboniferous, Ohio became a more Ecoregion#Terrestrial, terrestrial environment with an increased diversity of plants that formed expansive swampy River delta, deltas. Amphibians and reptiles began to inhabit the state at this time, and remained present into the ensuing Permian. A gap in the local rock record spans from this point until the start of the Pleistoce ...
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Fossil Taxa Described In 1878
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, Seashell, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in #Resin, amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock stratum, strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitativ ...
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Placoderms Of North America
Placodermi (from Greek πλάξ 'plate' and δέρμα 'skin', literally 'plate-skinned') is a class of armoured prehistoric fish, known from fossils, which lived from the Silurian to the end of the Devonian period. Their head and thorax were covered by articulated armoured plates and the rest of the body was scaled or naked, depending on the species. Placoderms were among the first jawed fish; their jaws likely evolved from the first of their gill arches. Placoderms are thought to be paraphyletic, consisting of several distinct outgroups or sister taxa to all living jawed vertebrates, which originated among their ranks. In contrast, one 2016 analysis concluded that placodermi are likely monophyletic, though these analyses have been further dismissed with more transitional taxa between placoderms and modern gnathosthomes, solidifying their paraphyletic status. Placoderms were also the first fish to develop pelvic fins, the precursor to hindlimbs in tetrapods, as well as true t ...
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Animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motility, able to move, can Sexual reproduction, reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of Cell (biology), cells, the blastula, during Embryogenesis, embryonic development. Over 1.5 million Extant taxon, living animal species have been Species description, described—of which around 1 million are Insecta, insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have Ecology, complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a Symmetry in biology#Bilate ...
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