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Dinocephalian Genera
Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included herbivorous, carnivorous, and omnivorous forms. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania. Description Apart from the Biarmosuchians, the dinocephalians are the least advanced therapsids, although still uniquely specialised in their own way. They retain a number of primitive characteristics (e.g. no secondary palate, small dentary) shared with their pelycosaur ancestors, although they are also more advanced in possessing therapsid adaptations like the expansion of the ilium and more erect limbs. They include carnivoro ...
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Early Permian
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Secondary Palate
The secondary palate is an anatomical structure that divides the nasal cavity from the oral cavity in many vertebrates. In human embryology, it refers to that portion of the hard palate that is formed by the growth of the two palatine shelves medially and their mutual fusion in the midline. It forms the majority of the adult palate and meets the primary palate at the incisive foramen. Clinical significance Secondary palate development begins in the sixth week of pregnancy and can lead to cleft palate when development goes awry. There are three major mechanisms known to cause this failure: #Growth retardation — Palatal shelves do not grow enough to meet each other. #Mechanical obstruction — Improper mouth size, or abnormal anatomical structures in the embryonic mouth prevent fully grown shelves from meeting each other. #Midline epithelial dysfunction (MED)Dudas et al. (2007): Palatal fusion – Where do the midline cells go? A review on cleft palate, a major hum ...
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Styracocephalus
''Styracocephalus platyrhynchus'' ('spike head') is an extinct species of tapinocephalian therapsids that lived during the Guadalupian epoch. ''Styracocephalus''s head ornament meant that it could be recognised from a distance. The most striking feature of ''Styracocephalus'' are the large backward-protruding tabular horns. The crest stuck upwards and backwards, but there is some variation in its shape, and this suggests that it changed throughout life and that it may be sexually dimorphic. ''Styracocephalus'' was a herbivore that may have been fully terrestrial or partly aquatic like the modern hippopotamus. It may have evolved from the estemmenosuchids. Its remains are known from South Africa but it probably had a wider distribution. It was around in length, with a , skull. See also * List of therapsids This list of therapsids is an attempt to create a comprehensive listing of all genera that have ever been included in the Therapsida excluding mammals and purely ve ...
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Estemmenosuchus
''Estemmenosuchus'' (meaning "crowned crocodile" in Greek) is an extinct genus of large, early omnivorous therapsid. It is believed and interpreted to have lived during the middle part of the Middle Permian around 267 million years ago. The two species, ''E. uralensis'' and ''E. mirabilis'', are characterised by distinctive horn-like structures, which were probably used for intra-specific display. Both species of ''Estemmenosuchus'' are from the Perm (or Cis-Urals) region of Russia. Two other estemmenosuchids, ''Anoplosuchus'' and ''Zopherosuchus'', are now considered females of the species ''E. uralensis''. There were many complete and incomplete skeletons found together. Description ''Estemmenosuchus'' could reach a body length of more than . Its skull was long and massive, up to in length, and possessed several sets of large horns, somewhat similar to the antlers of a moose, growing upward and outward from the sides and top of the head. The animal had a sprawling posture a ...
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Pachyostosis
Pachyostosis is a non-pathological condition in vertebrate animals in which the bones experience a thickening, generally caused by extra layers of lamellar bone. It often occurs together with bone densification (osteosclerosis), reducing inner cavities. This joint occurrence is called pachyosteosclerosis. However, especially in the older literature, "pachyostosis" is often used loosely, referring to all osseous specializations characterized by an increase in bone compactness and/or volume. It occurs in both terrestrial and, especially, aquatic or semi-aquatic vertebrates. In aquatic animals, such as seacows (manatees and dugongs), ''Thalassocnus'', and plesiosaurs, pachyostosis in the thoracic region provides (or provided) ballast against the air-filled lungs. This maintains neutral buoyancy in aquatic habitats. Most giant deer showed pronounced pachyostosis of the mandible and skull. It has been suggested that this served to store minerals for antler growth. Many Pachycephalosau ...
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Anteosaurus
''Anteosaurus'' (meaning "Antaeus reptile") is an extinct genus of large carnivorous dinocephalian synapsid. It lived at the end of the Guadalupian (= Middle Permian) during the Capitanian stage, about 265 to 260 million years ago in what is now South Africa. It is mainly known by cranial remains and few postcranial bones. With its skull reaching in length and a body size estimated at more than in length, and in weight, ''Anteosaurus'' was the largest known carnivorous non-mammalian synapsid and the largest terrestrial predator of the Permian period. Occupying the top of the food chain in the Middle Permian, its skull, jaws and teeth show adaptations to capture large prey like the giants titanosuchids and tapinocephalids dinocephalians and large pareiasaurs. As in many other dinocephalians the cranial bones of ''Anteosaurus'' are pachyostosed, but to a lesser extent than in tapinocephalid dinocephalians. In ''Anteosaurus'', pachyostosis mainly occurs in the form of horn-sha ...
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Titanosuchus
''Titanosuchus ferox'' ("Fierce titan crocodile") is an extinct species of dinocephalian therapsids that lived in the Middle Permian epoch in South Africa. Along with its close relatives, ''Jonkeria'' and ''Moschops'', ''Titanosuchus'' inhabited present-day South Africa around 265 million years ago, in the Late Permian. ''Titanosuchus'' was a carnivore which measured over 2.5 m long and might have eaten both ''Jonkeria'' and ''Moschops'', among other vertebrates. Its teeth included sharp incisors and fang-like canines, perfect for biting prey. ''Titanosuchus'' should not be confused with the therapsid ''Eotitanosuchus'', which belonged to a different family. ''Parascapanodon'' and ''Scapanodon'' were once thought to be distinct genera, but are now considered to be junior synonyms of ''Titanosuchus''.Boonstra, L. D., 1969, The fauna of the Tapinocephalus zone (Beaufort beds of the Karoo): Annals of the South African Museum, v. 56, part 1, p. 1-73. See also * List of therapsids ...
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Tapinocephalus
''Tapinocephalus'' ("humble head") is an extinct genus of large herbivorous dinocephalians that lived during the Middle Permian Period. These stocky, barrel-bodied animals were characterised by a massive bony skull roof and short weak snout. It is thought that, like the rest of the members of its family, the animals engaged in head-butting intraspecific behavior, possibly for territory or mates. The fossil remains (skull and postcranial elements) of ''Tapinocephalus'' are known from the Lower, Middle, and Upper part of the ''Tapinocephalus'' Assemblage Zone ( Capitanian age) of the Lower Beaufort Beds of the South African Karoo. Only the type species, ''T. atherstonei'' is now considered valid for this genus. In life, these animals were over in length and massed around , making them among the largest animals of their time. Classification ''Tapinocephalus atherstonei'' is known from a number of skulls and postcranial bones. The skull is large with a heavily pachyostotic ...
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Pareiasaur
Pareiasaurs (meaning "cheek lizards") are an extinct clade of large, herbivorous parareptiles. Members of the group were armoured with scutes which covered large areas of the body. They first appeared in southern Pangea during the Middle Permian, before becoming globally distributed during the Late Permian. Pareiasaurs were the largest reptiles of the Permian, reaching sizes equivalent to those of contemporary therapsids. Pareiasaurs became extinct at the end of the Permian during the Permian-Triassic extinction event. Description Pareiasaurs ranged in size from long, and may have weighed up to . They were stocky, with short tails, small heads, robust limbs, and broad feet. The cow-sized species '' Bunostegos'', which lived 260 million years ago, is the earliest known example of a tetrapod with a fully erect posture as its legs were positioned directly under its body. Pareiasaurs were protected by bony scutes called osteoderms that were set into the skin. Their heavy skulls wer ...
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Caseidae
Caseidae are an extinct family of basal synapsids that lived from the Late Carboniferous to Middle Permian between about 300 and 265 million years ago. Fossils of these animals come from the south-central part of the United States (Texas, Oklahoma, and Kansas), from various parts of Europe (European Russia, France, Germany, Sardinia, and Poland), and possibly from South Africa if the genus ''Eunotosaurus'' is indeed a caseid as some authors proposed in 2021. Caseids show great taxonomic and morphological diversity. The most basal taxa were small insectivorous and omnivorous forms that lived mainly in the Upper Carboniferous and Lower Permian, such as ''Eocasea'', ''Callibrachion'', and '' Martensius''. This type of caseid persists until the middle Permian with ''Phreatophasma'' and may be ''Eunotosaurus''. During the early Permian, the clade is mainly represented by many species that adopted a herbivorous diet. Some have evolved into gigantic forms that can reach in length, such ...
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Permian
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids ( reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their am ...
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Semiaquatic
In biology, semiaquatic can refer to various types of animals that spend part of their time in water, or plants that naturally grow partially submerged in water. Examples are given below. Semiaquatic animals Semiaquatic animals include: * Vertebrates ** Amphibious fish; also several types of normally fully aquatic fish such as the grunion and plainfin midshipman that spawn in the intertidal zone ** Some amphibians such as newts and salamanders, and some frogs such as fire-bellied toads and wood frogs. ** Some reptiles such as crocodilians, turtles, water snakes and marine iguanas. ** Penguins. ** Some rodents such as beavers, muskrats and capybaras. **Some insectivorous mammals such as desmans, water shrews and platypuses. ** Some carnivoran mammals, including seals, polar bears and otters. ** Hippopotamuses. * Semiterrestrial echinoderms of the intertidal zone, such as the "cliff-clinging" sea urchin ''Colobocentrotus atratus'' and the starfish '' Pisaster ochraceus'' ...
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