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Dennstaedtiineae
Dennstaedtiaceae is one of fifteen families in the order Polypodiales, the most derived families within fern, monilophytes (ferns). It comprises 10 genera with ca 240 known species, including one of the world's most abundant ferns, ''Pteridium aquilinum'' (bracken). Members of the order generally have large, highly divided leaves and have either small, round intramarginal sori with cup-shaped indusium, indusia (e.g. ''Dennstaedtia'') or linear marginal sori with a false indusium formed from the reflexed leaf margin (e.g. ''Pteridium''). The morphological diversity among members of the order has confused past taxonomy, but recent molecular studies have supported the monophyletic, monophyly of the order and the family.Smith, A. R., K. M. Pryer, et al. (2006). "A classification for extant ferns." Taxon 55(3): 705-731 The reclassification of Dennstaedtiaceae and the rest of the monilophytes was published in 2006, so most of the available literature is not updated. Characteristics * T ...
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Polypodiales
The order Polypodiales encompasses the major lineages of polypod ferns, which comprise more than 80% of today's fern species. They are found in many parts of the world including tropical, semitropical and temperate areas. Description Polypodiales are unique in bearing sporangia with a vertical annulus interrupted by the stalk and stomium. These sporangial characters were used by Johann Jakob Bernhardi to define a group of ferns he called the "Cathetogyratae"; the Pteridophyte Phylogeny Group has suggested reviving this name as the informal term cathetogyrates, to replace the ambiguously circumscribed term "polypods" when referring to the Polypodiales. The sporangia are born on stalks 1–3 cells thick and are often long-stalked. (In contrast, the Hymenophyllales have a stalk composed of four rows of cells.) The sporangia do not reach maturity simultaneously. Many groups in the order lack indusia, but when present, they are attached either along the edge of the indusium or in its ...
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Saccolomataceae
Saccolomataceae is a family of ferns in the order Polypodiales with about 19 species. It has been formerly treated as part of the Dennstaedtiaceae, however it has been classified as its own family according to Smith et al. (2006) The genus ''Saccoloma'' has been classified to include '' Orthiopteris'', but the phylogeny of the group not yet fully understood. The family includes a dozen known species. Description Saccolomataceae generally have the following characteristics: Terrestrial; rhizomes short-creeping to erect and trunk-like; petioles each with an omega-shaped vascular strand; blades pinnate to decompound and lacking articulate hairs; veins free; sori terminal on the veins; indusia pouch- or cup-shaped. Taxonomy In 2016 the Pteridophyte Phylogeny Group placed it as the sole family in the Saccolomatineae, one of six suborders making up the Polypodiales order. These suborders are phylogenetically related as shown in this cladogram A cladogram (from Greek ''clad ...
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Dennstaedtiales
The order Polypodiales encompasses the major lineages of polypod ferns, which comprise more than 80% of today's fern species. They are found in many parts of the world including tropical, semitropical and temperate areas. Description Polypodiales are unique in bearing sporangia with a vertical annulus interrupted by the stalk and stomium. These sporangial characters were used by Johann Jakob Bernhardi to define a group of ferns he called the "Cathetogyratae"; the Pteridophyte Phylogeny Group has suggested reviving this name as the informal term cathetogyrates, to replace the ambiguously circumscribed term "polypods" when referring to the Polypodiales. The sporangia are born on stalks 1–3 cells thick and are often long-stalked. (In contrast, the Hymenophyllales have a stalk composed of four rows of cells.) The sporangia do not reach maturity simultaneously. Many groups in the order lack indusia, but when present, they are attached either along the edge of the indusium or in its ...
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Fern
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except the lycopods, and differ from mosses and other bryophytes by being vascular, i.e., having specialized tissues that conduct water and nutrients and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaves called megaphylls, that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns, and ophioglossoid ferns. Ferns first ...
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Dennstaedtiaceae
Dennstaedtiaceae is one of fifteen families in the order Polypodiales, the most derived families within monilophytes (ferns). It comprises 10 genera with ca 240 known species, including one of the world's most abundant ferns, ''Pteridium aquilinum'' (bracken). Members of the order generally have large, highly divided leaves and have either small, round intramarginal sori with cup-shaped indusia (e.g. ''Dennstaedtia'') or linear marginal sori with a false indusium formed from the reflexed leaf margin (e.g. ''Pteridium''). The morphological diversity among members of the order has confused past taxonomy, but recent molecular studies have supported the monophyly of the order and the family.Smith, A. R., K. M. Pryer, et al. (2006). "A classification for extant ferns." Taxon 55(3): 705-731 The reclassification of Dennstaedtiaceae and the rest of the monilophytes was published in 2006, so most of the available literature is not updated. Characteristics * Terrestrial or scrambling (sca ...
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Monachosoraceae
Dennstaedtiaceae is one of fifteen families in the order Polypodiales, the most derived families within monilophytes (ferns). It comprises 10 genera with ca 240 known species, including one of the world's most abundant ferns, ''Pteridium aquilinum'' (bracken). Members of the order generally have large, highly divided leaves and have either small, round intramarginal sori with cup-shaped indusia (e.g. ''Dennstaedtia'') or linear marginal sori with a false indusium formed from the reflexed leaf margin (e.g. ''Pteridium''). The morphological diversity among members of the order has confused past taxonomy, but recent molecular studies have supported the monophyly of the order and the family.Smith, A. R., K. M. Pryer, et al. (2006). "A classification for extant ferns." Taxon 55(3): 705-731 The reclassification of Dennstaedtiaceae and the rest of the monilophytes was published in 2006, so most of the available literature is not updated. Characteristics * Terrestrial or scrambling (sca ...
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Lindsaeaceae
Lindsaeaceae is a pantropical family of ferns in the order Polypodiales. It contains six or seven genera with about 220 known species, some of which also extend into the more temperate regions of eastern Asia, New Zealand, and South America.Lehtonen ''et al.'': Phylogenetics and classification of the pantropical fern family Lindsaeaceae
in the ''Botanical Journal of the Linnaen Society'' 2010


Description

Characteristics include: Rhizomes short to long creeping; rhizomes with nonclathrate scales or uniseriate hairs; blades 1-3 pinnate or more divided; veins usually free; sori marginal or submarginal;

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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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Burmese Amber
Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The amber is of significant palaeontological interest due to the diversity of flora and fauna contained as inclusions, particularly arthropods including insects and arachnids but also birds, lizards, snakes, frogs and fragmentary dinosaur remains. The amber has been known and commercially exploited since the first century AD, and has been known to science since the mid-nineteenth century. Research on the deposit has attracted controversy due to its alleged role in funding internal conflict in Myanmar and hazardous working conditions in the mines where it is collected. Geological context, depositional environment and age The amber is found within the Hukawng Basin, a large Cretaceous-Cenozoic sedimentary basin within northern Myanmar. The s ...
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Cenomanian
The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the stratigraphic column deposited during the corresponding age. Both age and stage bear the same name. As a unit of geologic time measure, the Cenomanian Age spans the time between 100.5 and 93.9 million years ago (Mya). In the geologic timescale, it is preceded by the Albian and is followed by the Turonian. The Upper Cenomanian starts around at 95 Mya. The Cenomanian is coeval with the Woodbinian of the regional timescale of the Gulf of Mexico and the early part of the Eaglefordian of the regional timescale of the East Coast of the United States. At the end of the Cenomanian, an anoxic event took place, called the Cenomanian-Turonian boundary event or the "Bonarelli event", that is associated with a minor extinction event for marine spec ...
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Monophyletic
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic groups are typically characterised by shared derived characteristics ( synapomorphies), which distinguish organisms in the clade from other organisms. An equivalent term is holophyly. The word "mono-phyly" means "one-tribe" in Greek. Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic group'' consists of all of the descendants of a common ancestor minus one or more monophyletic groups. A '' polyphyletic group'' is characterized by convergent features or habits of scientific interest (for example, night-active primates, fruit trees, aquatic insects). The features by which a polyphyletic group is differentiated from others are not inherited from a common ancestor. These definitions have tak ...
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