Claosaurus Agilis
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Claosaurus Agilis
''Claosaurus'' ( ; Ancient Greek, Greek κλάω, ''klao'' meaning 'broken' and , ''sauros'' meaning 'lizard'; "broken lizard", referring to the odd position of the fossils when discovered) is a genus of hadrosauroid dinosaur that lived during the Late Cretaceous Period (geology), Period (Santonian-Campanian). Traditionally classified as an early member of the family Hadrosauridae, a 2008 analysis found ''Claosaurus agilis'' to be outside of the clade containing ''Hadrosaurus'' and other hadrosaurids, making it the closest non-hadrosaurid relative of true hadrosaurids within the clade Hadrosauria. Description Because of the insufficient fossil remains ("minority of skull and partial skeleton"), the size of an adult ''Claosaurus'' remains uncertain. However, Thomas Holtz gave a length estimate of and a mass estimate as that of a lion. Like other hadrosaurs, it was an herbivore. History Evidence of its existence was first found in the Niobrara Formation near the Smoky Hill ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Ant ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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Barnum Brown
Barnum Brown (February 12, 1873 – February 5, 1963), commonly referred to as Mr. Bones, was an American paleontologist. Named after the circus showman P. T. Barnum, he discovered the first documented remains of ''Tyrannosaurus'' during a career that made him one of the most famous fossil hunters working from the late Victorian era into the early 20th century. Fossil dinosaur expeditions Sponsored by the American Museum of Natural History (AMNH), Brown traversed the country bargaining and trading for fossils. His field was not limited to dinosaurs. He was known to collect or obtain anything of possible scientific value. Often, he simply sent money to have fossils shipped to the AMNH, and any new specimen of interest often resulted in a flurry of letters between the discoverer and Brown. After working a handful of years in Wyoming for AMNH in the late 1890s, Brown led an expedition to the Hell Creek Formation of southeastern Montana. There, in 1902, he discovered and exca ...
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Gastrolith
A gastrolith, also called a stomach stone or gizzard stone, is a rock held inside a gastrointestinal tract. Gastroliths in some species are retained in the muscular gizzard and used to grind food in animals lacking suitable grinding teeth. In other species the rocks are ingested and pass through the digestive system and are frequently replaced. The grain size depends upon the size of the animal and the gastrolith's role in digestion. Other species use gastroliths as ballast.Rondeau, et aLarval Anurans Adjust Buoyancy in Response to Substrate IngestionCopeia: February 2005, Vol. 2005, No. 1, pp. 188-195. Particles ranging in size from sand to cobble have been documented. Etymology Gastrolith comes from the Greek γαστήρ (''gastēr''), meaning "stomach", and λίθος (''lithos''), meaning "stone". Occurrence Among living vertebrates, gastroliths are common among crocodiles, alligators, herbivorous birds, seals and sea lions. Domestic fowl require access to ''grit''. Stone ...
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Nomen Dubium
In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a specimen belongs to that group or not. This may happen if the original type series (i. e. holotype, isotype, syntype or paratype) is lost or destroyed. The zoological and botanical codes allow for a new type specimen, or neotype, to be chosen in this case. A name may also be considered a ''nomen dubium'' if its name-bearing type is fragmentary or lacking important diagnostic features (this is often the case for species known only as fossils). To preserve stability of names, the ''International Code of Zoological Nomenclature'' allows a new type specimen, or neotype, to be chosen for a ''nomen dubium'' in this case. 75.5. Replacement of unidentifiable name-bearing type by a neotype. When an author considers that the taxonomic identity of a ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily "typ ...
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Archelon
''Archelon'' is an extinct marine turtle from the Late Cretaceous, and is the largest turtle ever to have been documented, with the biggest specimen measuring from head to tail and in body mass. It is known only from the Dakota Pierre Shale and has one species, ''A. ischyros''. In the past, the genus also contained ''A. marshii'' and ''A. copei'', though these have been reassigned to '' Protostega'' and ''Microstega'', respectively. The genus was named in 1895 by American paleontologist George Reber Wieland based on a skeleton from South Dakota, who placed it into the extinct family Protostegidae. The leatherback sea turtle (''Dermochelys coriacea'') was once thought to be its closest living relative, but now, Protostegidae is thought to be a completely separate lineage from any living sea turtle. ''Archelon'' had a leathery carapace instead of the hard shell seen in sea turtles. The carapace may have featured a row of small ridges, each peaking at in height. It had an esp ...
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South Dakota
South Dakota (; Sioux language, Sioux: , ) is a U.S. state in the West North Central states, North Central region of the United States. It is also part of the Great Plains. South Dakota is named after the Lakota people, Lakota and Dakota people, Dakota Sioux Native Americans in the United States, Native American tribes, who comprise a large portion of the population with nine Indian reservation, reservations currently in the state and have historically dominated the territory. South Dakota is the List of U.S. states and territories by area, seventeenth largest by area, but the List of U.S. states and territories by population, 5th least populous, and the List of U.S. states and territories by population density, 5th least densely populated of the List of U.S. states, 50 United States. As the southern part of the former Dakota Territory, South Dakota became a state on November 2, 1889, simultaneously with North Dakota. They are the 39th and 40th states admitted to the union; Pr ...
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Pierre Shale
The Pierre Shale is a geologic formation or series in the Upper Cretaceous which occurs east of the Rocky Mountains in the Great Plains, from Pembina Valley in Canada to New Mexico. The Pierre Shale was described by Meek and Hayden in 1862 in the Proceedings of the Academy of Sciences (Philadelphia). They described it as a dark-gray shale, fossiliferous, with veins and seams of gypsum, and concretions of iron oxide. The Pierre Shale is about 700 feet (210m) thick at the type locality. It overlies the Niobrara division and underlies the Fox Hills beds. It was named for an occurrence near Fort Pierre on the Missouri River in South Dakota. The Pierre Shale is of marine origin and was deposited in the Western Interior Seaway. It is correlative with other marine shales that occur farther west, such as the Bearpaw Shale, Mancos Shale and the Lewis Shale. It correlates with the Lea Park Formation in central Alberta. The Pierre is overlain by marginal marine deposits of the Fox H ...
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Edmontosaurus
''Edmontosaurus'' ( ) (meaning "lizard from Edmonton") is a genus of hadrosaurid (duck-billed) dinosaur. It contains two known species: ''Edmontosaurus regalis'' and ''Edmontosaurus annectens''. Fossils of ''E. regalis'' have been found in rocks of western North America that date from the late Campanian Stage (stratigraphy), stage of the Cretaceous Period (geology), Period 73 million years ago, while those of ''E. annectens'' were found in the same geographic region but in rocks dated to the end of the Maastrichtian stage of the Cretaceous, 66 million years ago. ''Edmontosaurus'' was one of the last non-bird, avian dinosaurs, and lived alongside dinosaurs like ''Triceratops'', ''Tyrannosaurus'', ''Ankylosaurus'', and ''Pachycephalosaurus'' shortly before the Cretaceous–Paleogene extinction event. ''Edmontosaurus'' included some of the largest hadrosaurid species, with ''E. annectens'' measuring up to in length and weighing around in average asymptotic body mass, althoug ...
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Kansas
Kansas () is a state in the Midwestern United States. Its capital is Topeka, and its largest city is Wichita. Kansas is a landlocked state bordered by Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to the west. Kansas is named after the Kansas River, which in turn was named after the Kansa Native Americans who lived along its banks. The tribe's name (natively ') is often said to mean "people of the (south) wind" although this was probably not the term's original meaning. For thousands of years, what is now Kansas was home to numerous and diverse Native American tribes. Tribes in the eastern part of the state generally lived in villages along the river valleys. Tribes in the western part of the state were semi-nomadic and hunted large herds of bison. The first Euro-American settlement in Kansas occurred in 1827 at Fort Leavenworth. The pace of settlement accelerated in the 1850s, in the midst of political wars over the slavery debate. Wh ...
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Smoky Hill River
The Smoky Hill River is a river in the central Great Plains of North America, running through Colorado and Kansas.Smoky Hill River. (2009). In Encyclopædia Britannica. Retrieved July 22, 2009, from Encyclopædia Britannica Online: http://www.britannica.com/EBchecked/topic/550068/Smoky-Hill-River Names The Smoky Hill gets its name from the Smoky Hills region of north-central Kansas through which it flows. Indigenous peoples of the Americas, American Indians living along the Smoky Hill considered it and the Kansas River to be the same river, and their names for it included Chetolah and Okesee-sebo. Early maps of European explorers called the river (also in combination with the Kansas River) the River of the Padoucas as its source is located in what was then Padouca (Comanche) territory. The USGS lists a number of other variant names for the Smoky Hill River, including Chitolah River, Fork of the Hill Buckaneuse, La Fourche de la Cote Boucaniere, La Touche de la Cote Bucanieus, M ...
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