Chaetothyriomycetes
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Chaetothyriomycetes
''Chaetothyriomycetidae'' is a subclass of fungi within the Eurotiomycetes. They are collectively termed the black yeast “Black yeasts”, sometimes also black fungi, dematiaceous fungi, microcolonial fungi or meristematic fungi is a diverse group of slow-growing microfungi which reproduce mostly asexually ( fungi imperfecti). Only few genera reproduce by budding ...s. References * Geiser DM, Gueidan C, Miadlikowska J, Lutzoni F, Kauff F, Hofstetter V, Fraker E, Schoch CL, Tibell L, Untereiner WA, Aptroot A, 2006. Eurotiomycetes: Eurotiomycetidae and Chaetothyriomycetidae. Mycologia 98, 1053–1064. Eurotiomycetes {{dothideomycetes-stub ...
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Fungus
A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from the other eukaryotic kingdoms, which by one traditional classification include Plantae, Animalia, Protozoa, and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single group of related organisms, named the ''Eumycota'' (''true f ...
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Ascomycota
Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The defining feature of this fungal group is the " ascus" (), a microscopic sexual structure in which nonmotile spores, called ascospores, are formed. However, some species of the Ascomycota are asexual, meaning that they do not have a sexual cycle and thus do not form asci or ascospores. Familiar examples of sac fungi include morels, truffles, brewers' and bakers' yeast, dead man's fingers, and cup fungi. The fungal symbionts in the majority of lichens (loosely termed "ascolichens") such as ''Cladonia'' belong to the Ascomycota. Ascomycota is a monophyletic group (it contains all descendants of one common ancestor). Previously placed in the Deuteromycota along with asexual species from other fungal taxa, asexual (or anamorphic) ascomyce ...
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Pezizomycotina
Pezizomycotina make up most of the Ascomycota fungi and include most lichenized fungi too. Pezizomycotina contains the filamentous ascomycetes and is a subdivision of the Ascomycota (fungi that form their spores in a sac-like ''ascus''). It is more or less synonymous with the older taxon Euascomycota. These fungi reproduce by fission rather than budding and this subdivision includes almost all the ascus fungi that have fruiting bodies visible to the naked eye (exception: genus ''Neolecta'', which belongs to the Taphrinomycotina). See the taxobox for a list of the classes that make up the Pezizomycotina. The old class Loculoascomycetes (consisting of all the bitunicate Ascomycota) has been replaced by the two classes Eurotiomycetes and Dothideomycetes. The rest of the Pezizomycotina also include the previously defined hymenial groups Discomycetes (now Leotiomycetes) and Pyrenomycetes (Sordariomycetes). Some important groups in Pezizomycotina include: Pezizomycetes (the opercula ...
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Eurotiomycetes
Eurotiomycetes is a large class of ascomycetes with cleistothecial ascocarps within the subphylum Pezizomycotina, currently containing around 3810 species according to the Catalogue of Life. It is the third largest lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship. It contains most of the fungi previously known morphologically as "Plectomycetes".


Systematics and phylogeny


Internal relationships

The class Eurotiomycetes was circumscription (taxonomy), circumscribed in 1997 by Sweden, Swedish mycologists Ove Erik Eriksson and Katarina Winka. At that time ...

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Arthoniales
The Arthoniales is the second largest order of mainly crustose lichens, but fruticose lichens are present as well. The order contains around 1500 species, while the largest order with lichenized fungi, the Lecanorales, contains more than 14000 species. Classification The Arthoniales is one of two orders of the class Arthoniomycetes within the phylum Ascomycota. The order includes seven families ( Andreiomycetacae, Arthoniaceae, Chrysotrichaceae, Lecanographaceae, Opegraphaceae, Roccellaceae and Roccellographaceae). Lecanographaceae, Roccellographaceae, Opegraphaceae and Roccellaceae are well-supported families within Arthoniales, and they were circumscribed in 2011. Andreiomycetaceae was described as a new family by Hodkinson and Lendemer in 2013. The Arthoniales is the sister group to Dothideomycetes. Figure 1. Cladogram of the Arthoniales, rooted with ''Curvularia brachyspora'', ''Cudonia circinans'' and ''Seynesia erumpens'' as the outgroup. The cladogram shows the ...
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Chaetothyriales
The Chaetothyriales are an order of ascomycetous fungi in the class Eurotiomycetes and within the subclass Chaetothyriomycetidae. The order was circumscribed in 1987 by mycologist Margaret Elizabeth Barr-Bigelow. Families and genera , Species Fungorum includes 9 families, 97 genera, and 691 species in the Chaetothyriales. The following list shows the families, genera, and number of species in the Chaetothyriales, adapted from a recent (2020) taxonomic and nomenclatural review of the order. * Chaetothyriaceae ::'' Actinocymbe'' – 3 spp. ::'' Aithaloderma'' – 12 spp. ::'' Aphanophora'' – 1 sp. ::'' Arthrophiala'' – 1 sp. ::'' Camptophora'' – 2 spp. ::'' Ceramothyrium'' – 39 spp. ::'' Ceratocarpia'' – 3 spp. ::'' Chaetothyriomyces'' – 1 sp. ::'' Chaetothyrium'' – 67 spp. ::'' Cyphellophoriella'' – 1 sp. ::'' Euceramia'' – 3 spp. ::'' Longihyalospora'' – 2 spp. ::'' Microcallis'' – 9spp. ::'' Nullicamyces'' – 1 sp. ::'' Phaeosaccardinula'' – 41 spp. :: ...
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Black Yeast
“Black yeasts”, sometimes also black fungi, dematiaceous fungi, microcolonial fungi or meristematic fungi is a diverse group of slow-growing microfungi which reproduce mostly asexually (fungi imperfecti). Only few genera reproduce by budding cells, while in others hyphal or meristematic (isodiametric) reproduction is preponderant. Black yeasts share some distinctive characteristics, in particular a dark colouration ( melanisation) of their cell wall. Morphological plasticity, incrustation of the cell wall with melanins and presence of other protective substances like carotenoids and mycosporines represent passive physiological adaptations which enable black fungi to be highly resistant against environmental stresses. The term " polyextremotolerance" has been introduced to describe this phenotype, a good example of which is the species ''Aureobasidium pullulans''. Presence of 1,8-dihydroxynaphthalene melanin in the cell wall confers to the microfungi their characteristic oli ...
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