Santalales
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Santalales
The Santalales are an order of flowering plants with a cosmopolitan distribution, but heavily concentrated in tropical and subtropical regions. It derives its name from its type genus ''Santalum'' (sandalwood). Mistletoe is the common name for a number of parasitic plants within the order. Overview Many of the members of the order are parasitic plants, mostly hemiparasites, able to produce sugars through photosynthesis, but tapping the stems or roots of other plants to obtain water and minerals; some (e.g. ''Arceuthobium'') are obligate parasites, have low concentrations of chlorophyll within their shoots (1/5 to 1/10 of that found in their host's foliage), and derive the majority of their sustenance from their hosts' vascular tissues (water, micro- and macronutrients, and sucrose). Most have seeds without testae (seed coats), which is unusual for flowering plants. Classification The APG IV system of 2016 includes seven families. As in the earlier APG III system, it was ...
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Olacaceae
Olacaceae is a family of flowering plants in the order Santalales. They are woody plants, native throughout the tropical regions of the world. , the circumscription of the family varies; some sources maintain a broad family, others split it into seven segregate families. Taxonomy The 1998 APG system and the 2003 APG II system assign it to the order Santalales in the clade core eudicots. Prior to the advent of molecular data, the circumscription of the family Olacaceae varied widely between different authorities.Valéry Malécot and Daniel L. Nickrent. 2008. "Molecular Phylogenetic Relationships of Olacaceae and Related Santalales". ''Systematic Botany'' 33(1):97-106. Among these various classifications, about 30 genera were included in the family. 15 genera are recognized for Olacaceae by the Germplasm Resources Information Network. The phylogenetic investigation published in 2008 recovered seven clades that were well-supported by molecular and morphological characters, but no for ...
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Loranthaceae
Loranthaceae, commonly known as the showy mistletoes, is a family of flowering plants. It consists of about 75 genera and 1,000 species of woody plants, many of them hemiparasites. The three terrestrial species are ''Nuytsia floribunda'' (the Western Australian Christmas tree), ''Atkinsonia ligustrina'' (from the Blue Mountains of Australia), and '' Gaiadendron punctatum'' (from Central/South America.) Loranthaceae are primarily xylem parasites, but their haustoria may sometimes tap the phloem, while '' Tristerix aphyllus'' is almost holoparasitic. For a more complete description of the Australian Loranthaceae, seFlora of Australia online, for the Malesian Loranthaceae seFlora of Malesia Originally, Loranthaceae contained all mistletoe species, but the mistletoes of Europe and North America (''Viscum'', ''Arceuthobium'', and ''Phoradendron'') belong to the family Santalaceae. The APG II system 2003 assigns the family to the order Santalales in the clade core eudicots. Phylog ...
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Mistletoe
Mistletoe is the common name for obligate hemiparasitic plants in the order Santalales. They are attached to their host tree or shrub by a structure called the haustorium, through which they extract water and nutrients from the host plant. The name mistletoe originally referred to the species ''Viscum album'' (European mistletoe, of the family Santalaceae in the order Santalales); it is the only species native to the British Isles and much of Europe. A related species with red rather than white fruits, ''Viscum cruciatum'', occurs in Southwest Spain and Southern Portugal, as well as in Morocco in North Africa and in southern Africa. The genus ''Viscum'' is not native to North America, but ''Viscum album'' was introduced to Northern California in 1900. The eastern mistletoe native to North America, ''Phoradendron leucarpum'', belongs to a distinct genus of the family Santalaceae. European mistletoe has smooth-edged, oval, evergreen leaves borne in pairs along the woody st ...
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Balanophoraceae
The Balanophoraceae are a subtropical to tropical family of obligate parasitic flowering plants, notable for their unusual development and formerly obscure affinities. In the broadest circumscription, the family consists of 16 genera. Alternatively, three genera may be split off into the segregate family Mystropetalaceae. The plants are normally found in moist inland forests growing on tree roots and have an above ground inflorescence with the overall appearance of a fungus, composed of numerous minute flowers. The inflorescences develop inside the tuberous underground part of the plant, before rupturing it and surfacing. The plants are monoecious, or dioecious, and the fruits are indehiscent drupes or nuts. The underground portion, which attaches itself to the host, looks like a tuber, and is not a proper root system. The plants contain no chlorophyll. ''Balanophora'' means "bearing an acorn" (from the shape of the female inflorescence). Taxonomy In the classification system of ...
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APG III System
The APG III system of flowering plant classification is the third version of a modern, mostly molecular-based, system of plant taxonomy being developed by the Angiosperm Phylogeny Group (APG). Published in 2009, it was superseded in 2016 by a further revision, the APG IV system. Along with the publication outlining the new system, there were two accompanying publications in the same issue of the Botanical Journal of the Linnean Society: * The first, by Chase & Reveal, was a formal phylogenetic classification of all land plants (embryophytes), compatible with the APG III classification. As the APG have chosen to eschew ranks above order, this paper was meant to fit the system into the existing Linnaean hierarchy for those that prefer such a classification. The result was that all land plants were placed in the class Equisetopsida, which was then divided into 16 subclasses and a multitude of superorders. * The second, by Haston ''et al.'', was a linear sequence of families followi ...
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Misodendraceae
''Misodendrum'' is a genus of hemiparasites which grow as mistletoes on various species of ''Nothofagus''. Its species are all restricted to South America. The name of the genus is incorrectly spelt in a number of ways, including ''Misodendron'' and ''Myzodendron''. ''Misodendrum'' is placed in its own family, Misodendraceae, in the order Santalales. These plants have the common name of feathery mistletoes. Species , The Plant List accepts the following species: *''Misodendrum angulatum'' Phil. *'' Misodendrum brachystachyum'' DC. *'' Misodendrum gayanum'' Tiegh. *'' Misodendrum linearifolium'' DC. *'' Misodendrum macrolepis'' Phil. *'' Misodendrum oblongifolium'' DC. *'' Misodendrum punctulatum'' Banks ex DC. *'' Misodendrum quadriflorum'' DC. See also *Parasitic plant A parasitic plant is a plant that derives some or all of its nutritional requirements from another living plant. They make up about 1% of angiosperms and are found in almost every biome. All parasitic plants ...
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Santalaceae
The Santalaceae, sandalwoods, are a widely distributed family of flowering plants (including small trees, shrubs, perennial herbs, and epiphytic climbersHewson & George t al.br>''Santalaceae'' taxonomy, 1984, pp. 191-194.) which, like other members of Santalales, are partially parasitic on other plants. Its flowers are bisexual or, by abortion ("flower drop"), unisexual.Pilger, R''Santalaceae''(with 17 figures). R. Br. Prodr. Fl. Nov. Holl. (1810) 350, pp. 1-45. Modern treatments of the Santalaceae include the family Viscaceae (mistletoes), previously considered distinct. The APG II system of 2003 recognises the family and assigns it to the order Santalales in the clade core eudicots. However, the circumscription by APG is much wider than accepted by previous classifications, including the plants earlier treated in families Eremolepidaceae and Viscaceae. It includes about 1,000 species in 43 genera. Many have reported traditional and cultural uses, including as medicine. Genera ...
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Opiliaceae
Opiliaceae is a family of flowering plants comprising 11 genera and 33 known species. It consists of tropical woody plants. Several genera contain parasitic species. The biggest genus, in number of species and in stature of the individual plants, is ''Agonandra'', the only American genus. Since it was first described this family has been recognized universally by taxonomists. The APG II system, of 2003 (unchanged from the APG system, of 1998), also recognizes this family and assigns it to the order Santalales in the clade core eudicots The eudicots, Eudicotidae, or eudicotyledons are a clade of flowering plants mainly characterized by having two seed leaves upon germination. The term derives from Dicotyledons. Traditionally they were called tricolpates or non-magnoliid dicots .... References External links Opiliaceae at Parasitic Plants Eudicot families {{Santalales-stub ...
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APG IV System
The APG IV system of flowering plant classification is the fourth version of a modern, mostly molecular-based, system of plant taxonomy for flowering plants (angiosperms) being developed by the Angiosperm Phylogeny Group (APG). It was published in 2016, seven years after its predecessor the APG III system was published in 2009, and 18 years after the first APG system was published in 1998. In 2009, a linear arrangement of the system was published separately; the APG IV paper includes such an arrangement, cross-referenced to the 2009 one. Compared to the APG III system, the APG IV system recognizes five new orders (Boraginales, Dilleniales, Icacinales, Metteniusales and Vahliales), along with some new families, making a total of 64 angiosperm orders and 416 families. In general, the authors describe their philosophy as "conservative", based on making changes from APG III only where "a well-supported need" has been demonstrated. This has sometimes resulted in placements that a ...
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Schoepfiaceae
Schoepfiaceae is a family of flowering plants recognized in the APG III system of 2009. The family was previously only recognized by few taxonomists; the plants in question usually being assigned to family Olacaceae and Santalaceae. The genus ''Schoepfia'' is, according to molecular studies, more closely related to the families Misodendraceae and Loranthaceae, and to uphold the criteria of monophyly it must be excluded from Olacaceae. Further investigations have also shown that the South American genera ''Arjona'' and '' Quinchamalium'' (previously in the family Santalaceae) are part of this family. Thus the family contains three genera. ''Quinchamalium'' was thought to comprise 21 species in 2015, but this was reduced to a single widespread and morphologically variable species in that year. Christenhusz ''et al''. claimed there was a total of 58 known species in 2016, but for some reason counted 25 species of ''Quinchamalium'' and eight of ''Arjona''. *''Arjona'' Cavanilles (5 o ...
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Parasitic Plant
A parasitic plant is a plant that derives some or all of its nutritional requirements from another living plant. They make up about 1% of angiosperms and are found in almost every biome. All parasitic plants develop a specialized organ called the haustorium, which penetrates the host plant, connecting them to the host vasculature – either the xylem, phloem, or both. For example, plants like ''Striga'' or ''Rhinanthus'' connect only to the xylem, via xylem bridges (xylem-feeding). Alternately, plants like ''Cuscuta'' and some members of ''Orobanche'' connect to both the xylem and phloem of the host. This provides them with the ability to extract water and nutrients from the host. Parasitic plants are classified depending on the location where the parasitic plant latches onto the host (root or stem), the amount of nutrients it requires, and their photosynthetic capability. Some parasitic plants can locate their host plants by detecting volatile chemicals in the air or soil given ...
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Santalum
''Santalum'' is a genus of woody flowering plants in the Santalaceae family, the best known and commercially valuable of which is the Indian sandalwood tree, '' S. album''. Members of the genus are trees or shrubs. Most are root parasites which photosynthesize their own food, but tap the roots of other species for water and inorganic nutrients. Several species, most notably ''S. album'', produce highly aromatic wood, used for scents and perfumes and for herbal medicine. About 25 known species range across the Indomalayan, Australasian, and Oceanian realms, from India through Malesia to the Pacific Islands, as far as Hawaii and the Juan Fernández Islands off the coast of South America. Indian sandalwood (''S. album'') is found in the tropical dry deciduous forests of India, the Lesser Sunda Islands of Indonesia, and Arnhem Land of northern Australia. It is the only species of the genus found on the Asian mainland, and may have been introduced to India from the Less ...
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