Rhynchonellata
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Rhynchonellata
The Rhynchonellata is a class of Lower Cambrian to Recent articulate brachiopods that combines orders from within the Rhynchonelliformea (Articulata revised) with well developed pedicle attachment. Shell forms vary from those with wide hinge lines to beaked forms with virtually no hinge line and from generally smooth to strongly plicate. Most all are biconvex. Lophophores vary and include both looped and spiraled forms. Although morphologically distinct, included orders follow a consistent phylogenetic sequence. Orders Orders assigned to the Rhynchonellata, in temporal sequence, include: * Protorthida ''L Camb - U Dev'' * Orthida ''L Camb - U Perm'' * Pentamerida ''M Camb - U Dev'' * Atrypida ''L Ord - U Dev'' * Rhynchonellida ''L Ord -Rec'' * Spiriferida ''U Ord - U Perm'' * Athyridida ''U Ord - L Jura'' * Spiriferinida ''L Dev - L Jura'' * Terebratulida ''L Dev - Rec'' * Thecideida ''U Trias - Rec'' The Atrypida, Athyridida, Spiriferida, and Spiriferinida were pr ...
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Rhynchonellata
The Rhynchonellata is a class of Lower Cambrian to Recent articulate brachiopods that combines orders from within the Rhynchonelliformea (Articulata revised) with well developed pedicle attachment. Shell forms vary from those with wide hinge lines to beaked forms with virtually no hinge line and from generally smooth to strongly plicate. Most all are biconvex. Lophophores vary and include both looped and spiraled forms. Although morphologically distinct, included orders follow a consistent phylogenetic sequence. Orders Orders assigned to the Rhynchonellata, in temporal sequence, include: * Protorthida ''L Camb - U Dev'' * Orthida ''L Camb - U Perm'' * Pentamerida ''M Camb - U Dev'' * Atrypida ''L Ord - U Dev'' * Rhynchonellida ''L Ord -Rec'' * Spiriferida ''U Ord - U Perm'' * Athyridida ''U Ord - L Jura'' * Spiriferinida ''L Dev - L Jura'' * Terebratulida ''L Dev - Rec'' * Thecideida ''U Trias - Rec'' The Atrypida, Athyridida, Spiriferida, and Spiriferinida were pr ...
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Articulata (Brachiopoda)
Rhynchonelliformea is a major subphylum and clade of brachiopods. It is equivalent to the former class Articulata, which was used previously in brachiopod taxonomy. Articulate brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations (hence the name) and separate sets of simple opening and closing muscles. The name, Rhynchoelliformea, which replaces Articulata, which is also a class of crinoids, comes from the family Rhynchonellidae, which however is no more representative of articulate brachiopods than any other group such as spirifers or strophomenids. It just happens to be the name chosen, based on an included taxon. The main difference between the Rhynchonelliformea described in the Treatise Part H, revised 2000/2007, and the Articulata of the Treatise part H, 1965, lies in the groups included, their taxonomic po ...
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Rhynchonelliformea
Rhynchonelliformea is a major subphylum and clade of brachiopods. It is equivalent to the former class Articulata, which was used previously in brachiopod taxonomy. Articulate brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations (hence the name) and separate sets of simple opening and closing muscles. The name, Rhynchoelliformea, which replaces Articulata, which is also a class of crinoids, comes from the family Rhynchonellidae, which however is no more representative of articulate brachiopods than any other group such as spirifers or strophomenids. It just happens to be the name chosen, based on an included taxon. The main difference between the Rhynchonelliformea described in the Treatise Part H, revised 2000/2007, and the Articulata of the Treatise part H, 1965, lies in the groups included, their taxonomic po ...
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Athyridida
Athyridida is an order of Paleozoic brachiopods included in the Rhynchonellata, which makes up part of the articulate brachiopods. The Athyridida are the Rostrospracea of R.C Moore, 1952,Moore, Lalcker and Fischer, 1952, Invertebrate Fossils, McGraw-Hill considered at that time to be a suborder of the Spiriferida. As with the Spiriferida, the Athyridida have outwardly directed spiral brachidia that support the lophophores on either side, but instead have non-plicate shells with rounded outlines and prominent beaks but almost no inner areas on the pedicle valve. Athyridids began early in the Silurian, reached their greatest diversity in the following Devonian, and from then declined steadily until almost becoming extinct at the end of the Permian The order rejuvenated somewhat during the Triassic, only to decline again until becoming extinct in the Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (strat ...
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Pentamerida
Pentamerida is an order of biconvex, impunctate shelled, articulate brachiopods that are found in marine sedimentary rocks that range from the Middle Cambrian through the Devonian.Moore, Lalcker and Fischer, 1952, Invertebrate Fossils, McGraw-Hill Pentamerids are characterized by a short hinge line where the two valves articulate, inner areas above the hinge line that slope inwardly from the beak of each valve, and a well-developed spondylium on the pedicle valve. The spondylium is a raised platform for muscle attachment found in the middle of the interior pedicle valve, generally toward the hinge and beak. The pedicle valve is the one that the pedicle, or hold-fast stalk, attaches to. The brachidia, which hold the lophophore, the ciliated feeding arms, are looped, as in the Orthida. The Short hinge line helps distinguish the pentamerids from the ancestral orthids from which they are obviously derived. The hinge line is not as short as found in the Rhynchonellida or Athyridida ...
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Brachiopod
Brachiopods (), phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major categories are traditionally recognized, articulate and inarticulate brachiopods. The word "articulate" is used to describe the tooth-and-groove structures of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic skeletal feature, by which the two main groups can be readily distinguished as fossils. Articulate brachiopods have toothed hinges and simple, vertically-oriented opening and closing muscles. Conversely, inarticulate brachiopods have weak, untoothed hinges and a more complex system of vertical and oblique (diagonal) muscles used to keep the two valves aligned. In many brachiopods, a ...
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Protorthida
Protorthida is an extinct order of Rhynchonellate brachiopods containing the taxa: * Superfamily Protorthoidea ** Family Protorthidae ** Family Arctohedridae ** Family Leioriidae * Superfamily Skenidioidea ** Family Skenidiidae References Rhynchonellata {{brachiopod-stub ...
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Orthida
Orthida is an extinct order of brachiopods which appeared during the Early Cambrian period and became very diverse by the Ordovician, living in shallow-shelf seas. Orthids are the oldest member of the subphylum Rhynchonelliformea, and is the order from which all other brachiopods of this group stem. Physically they are usually strophic, with well-developed interareas. They also commonly have radiating ribs, sulcus, and fold structures. Typically one valve, often the brachial valve, is flatter than the other. The interior structure of the brachial valves are usually simple. In shape they are sub-circular to elliptical, with typically biconvex valves. There is some debate over the forms that first appeared of this order as to how they should be classified. However, they began to differentiate themselves by the late Early Cambrian period, and by the late Cambrian period had diversified into numerous varieties and reach 2 to 5 cm in width. Specimens from the late Cambria ...
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Thecideida
Thecideida is an order of cryptic articulate brachiopods characterized by their small size and habit of cementing their ventral valves to hard substrates such as shells, rocks and carbonate hardgrounds. Thecideides first appear in the Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ... (Jaecks and Carlson, 2001) and are common today (Lüter, 2005; Lüter ''et al.'', 2007). Taxonomy Order Thecideida *Superfamily Thecideoidea Gray, 1840 *Superfamily Thecospiroidea Bittner, 1890 References Paleobiology Database on thecideides* * * Brachiopod orders Rhynchonellata {{Brachiopod-stub ...
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Treatise On Invertebrate Paleontology
The ''Treatise on Invertebrate Paleontology'' (or ''TIP'') published by the Geological Society of America and the University of Kansas Press, is a definitive multi-authored work of some 50 volumes, written by more than 300 paleontologists, and covering every phylum, class, order, family, and genus of fossil and extant (still living) invertebrate animals. The prehistoric invertebrates are described as to their taxonomy, morphology, paleoecology, stratigraphic and paleogeographic range. However, taxa with no fossil record whatsoever have just a very brief listing. Publication of the decades-long ''Treatise on Invertebrate Paleontology'' is a work-in-progress; and therefore it is not yet complete: For example, there is no volume yet published regarding the post-Paleozoic era caenogastropods (a molluscan group including the whelk and Common periwinkle, periwinkle). Furthermore, every so often, previously published volumes of the ''Treatise'' are revised. Evolution of the proje ...
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Terebratulida
Terebratulids are one of only three living orders of articulate brachiopods, the others being the Rhynchonellida and the Thecideida. Craniida and Lingulida include living brachiopods, but are inarticulates. The name, Terebratula, may be derived from the Latin "terebra", meaning "hole-borer". The perceived resemblance of terebratulid shells to ancient Roman oil lamps gave the brachiopods their common name "lamp shell". Terebratulids typically have biconvex shells that are usually ovoid to circular in outline. They can be either smooth or have radial ribbing. The lophophore support is loop shaped in contrast to the spiralia of similar looking spiriferids. Terebratulids are also distinguished by a very short hinge line, and the shell is punctate in microstructure. There is a circular pedicle opening, or foramen, located in the beak. Terebratulids may have evolved from Atrypids during the early or Middle Silurian. Early genera were almost circular to elongate-oval, with smooth or ...
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Lower Cambrian
The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Cambrian biolo ...
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