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Rhynchonelliformea
Rhynchonelliformea is a major subphylum and clade of brachiopods. It is equivalent to the former class Articulata, which was used previously in brachiopod taxonomy. Articulate brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations (hence the name) and separate sets of simple opening and closing muscles. The name, Rhynchoelliformea, which replaces Articulata, which is also a class of crinoids, comes from the family Rhynchonellidae, which however is no more representative of articulate brachiopods than any other group such as spirifers or strophomenids. It just happens to be the name chosen, based on an included taxon. The main difference between the Rhynchonelliformea described in the Treatise Part H, revised 2000/2007, and the Articulata of the Treatise part H, 1965, lies in the groups included, their taxonomic po ...
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Brachiopod
Brachiopods (), phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major categories are traditionally recognized, articulate and inarticulate brachiopods. The word "articulate" is used to describe the tooth-and-groove structures of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic skeletal feature, by which the two main groups can be readily distinguished as fossils. Articulate brachiopods have toothed hinges and simple, vertically-oriented opening and closing muscles. Conversely, inarticulate brachiopods have weak, untoothed hinges and a more complex system of vertical and oblique (diagonal) muscles used to keep the two valves aligned. In many brachiopods, a ...
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Treatise On Invertebrate Paleontology
The ''Treatise on Invertebrate Paleontology'' (or ''TIP'') published by the Geological Society of America and the University of Kansas Press, is a definitive multi-authored work of some 50 volumes, written by more than 300 paleontologists, and covering every phylum, class, order, family, and genus of fossil and extant (still living) invertebrate animals. The prehistoric invertebrates are described as to their taxonomy, morphology, paleoecology, stratigraphic and paleogeographic range. However, taxa with no fossil record whatsoever have just a very brief listing. Publication of the decades-long ''Treatise on Invertebrate Paleontology'' is a work-in-progress; and therefore it is not yet complete: For example, there is no volume yet published regarding the post-Paleozoic era caenogastropods (a molluscan group including the whelk and Common periwinkle, periwinkle). Furthermore, every so often, previously published volumes of the ''Treatise'' are revised. Evolution of the proje ...
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Craniformea
Craniata is a class of brachiopods originating in the Cambrian period and still extant today. It is the only class within the subphylum Craniiformea, one of three major subphyla of brachiopods alongside linguliforms and rhynchonelliforms. Craniata is divided into three orders: the extinct Craniopsida and Trimerellida, and the living Craniida, which provides most information on their biology. Living members of the class have shells which are composed of calcite, though some extinct forms my have aragonite shells. The shells are inarticulate (lack a hinge with distinct tooth-and-socket connections) and are usually rounded in outline. There is no pedicle, with the rear edge of the body cavity having the form of a smooth and flat wall perforated by the anus. This class of brachiopods has an unsupported lophophore with only a single row of tentacles. In the absence of a pedicle, the shell is usually attached directly to a hard substrate. Many craniiforms are encrusting animals whic ...
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Chileata
A rhynchonelliform Rhynchonelliformea is a major subphylum and clade of brachiopods. It is equivalent to the former class Articulata, which was used previously in brachiopod taxonomy. Articulate brachiopods have many anatomical differences relative to "inarticulate" ... class, characterised by the presence of a colleplax; closest relative (?): ''Salanygolina''.Holmer, L.E., Stolk, S.P., Skovsted, C.B., Balthasar, U., and Popov, L. (2009). The enigmatic early Cambrian ~Salanygolina~ - A stem group of rhynchonelliform chileate brachiopods? Palaeontology 52, 1–10. Salanygolina Thought to be a sister group. References Brachiopods {{paleo-protostome-stub ...
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Strophomenata
Strophomenata is an extinct class of brachiopods in the subphylum Rhynchonelliformea. Orders Billingsellida An order that contains the suborder Clitambonitidina (and others?) . Orthotetida An order or superfamily that includes the Chilidiopsoidea.LE Holmer, Z Zhang, TP Topper, and L Popov 2017. The attachment strategies of Cambrian kutorginate brachiopods: the curious case of two pedicle openings and their phylogenetic significance. Journal of Paleontology, 2017. Ontogeny given by Bassett and Popov (2017), resembling that of Kutorginides. Believed to be sister to Strophomenides, = Strophomenida + Billinsellida + Productida.Bassett, M.G., and Popov, L.E. (2017). Earliest ontogeny of the Silurian orthotetide brachiopod ~Coolinia~ and its significance for interpreting strophomenate phylogeny. Lethaia 50, 504–510. Includes the Chileids (e.g. ''Coolinia''). References External links

* * * Strophomenata, Prehistoric protostome classes {{paleo-protostome-stub ...
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Rhynchonellata
The Rhynchonellata is a class of Lower Cambrian to Recent articulate brachiopods that combines orders from within the Rhynchonelliformea (Articulata revised) with well developed pedicle attachment. Shell forms vary from those with wide hinge lines to beaked forms with virtually no hinge line and from generally smooth to strongly plicate. Most all are biconvex. Lophophores vary and include both looped and spiraled forms. Although morphologically distinct, included orders follow a consistent phylogenetic sequence. Orders Orders assigned to the Rhynchonellata, in temporal sequence, include: * Protorthida ''L Camb - U Dev'' * Orthida ''L Camb - U Perm'' * Pentamerida ''M Camb - U Dev'' * Atrypida ''L Ord - U Dev'' * Rhynchonellida ''L Ord -Rec'' * Spiriferida ''U Ord - U Perm'' * Athyridida ''U Ord - L Jura'' * Spiriferinida ''L Dev - L Jura'' * Terebratulida ''L Dev - Rec'' * Thecideida ''U Trias - Rec'' The Atrypida, Athyridida, Spiriferida, and Spiriferinida were pr ...
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Subphylum
In zoological nomenclature, a subphylum is a taxonomic rank below the rank of phylum. The taxonomic rank of "subdivision (rank), subdivision" in fungi and plant taxonomy is equivalent to "subphylum" in zoological taxonomy. Some plant taxonomists have also used the rank of subphylum, for instance monocotyledons as a subphylum of phylum Angiospermae and vertebrates as a subphylum of phylum Chordata. Taxonomic rank Subphylum is: #subordinate to the phylum #superordinate to the infraphylum. Where convenient, subphyla in turn can be divided into infraphyla; in turn such an infraphylum also would be superordinate to any classes or superclasses in the hierarchy. Examples Not all fauna phyla are divided into subphyla. Those that are include: *Arthropoda: divided into subphyla Trilobitomorpha, Chelicerata, Myriapoda, Hexapoda and Crustacean, Crustacea, *Brachiopoda: divided into subphyla Linguliformea, Craniformea and Rhynchonelliformea, *Chordata: divided into Tunicate, Tunicata, ...
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Phylogeny
A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological species or other entities based upon similarities and differences in their physical or genetic characteristics. All life on Earth is part of a single phylogenetic tree, indicating common ancestry. In a ''rooted'' phylogenetic tree, each node with descendants represents the inferred most recent common ancestor of those descendants, and the edge lengths in some trees may be interpreted as time estimates. Each node is called a taxonomic unit. Internal nodes are generally called hypothetical taxonomic units, as they cannot be directly observed. Trees are useful in fields of biology such as bioinformatics, systematics, and phylogenetics. ''Unrooted'' trees illustrate only the relatedness of the leaf nodes and do not require the ancestral root to be ...
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Athyridida
Athyridida is an order of Paleozoic brachiopods included in the Rhynchonellata, which makes up part of the articulate brachiopods. The Athyridida are the Rostrospracea of R.C Moore, 1952,Moore, Lalcker and Fischer, 1952, Invertebrate Fossils, McGraw-Hill considered at that time to be a suborder of the Spiriferida. As with the Spiriferida, the Athyridida have outwardly directed spiral brachidia that support the lophophores on either side, but instead have non-plicate shells with rounded outlines and prominent beaks but almost no inner areas on the pedicle valve. Athyridids began early in the Silurian, reached their greatest diversity in the following Devonian, and from then declined steadily until almost becoming extinct at the end of the Permian The order rejuvenated somewhat during the Triassic, only to decline again until becoming extinct in the Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (strat ...
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