Oncocerida
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Oncocerida
The Oncocerida comprise a diverse group of generally small nautiloid cephalopods known from the Middle Ordovician to the Mississippian (early Carboniferous; one possible member is known from the Early Permian), in which the connecting rings are thin and siphuncle segments are variably expanded (Flower, 1950). At present the order consists of some 16 families, a few of which, such as the Oncoceratidae, Brevicoceratidae, and Acleistoceratidae contain a fair number of genera each while others like the Trimeroceratidae and Archiacoceratidae are represented by only two or three (Sweet, 1964). Physical characteristics The shells of oncocerids are primarily somewhat compressed cyrtoconic brevicones. More advanced forms include gyrocones, serpenticones, torticones, and elongate orthocones and cyrtocones, reflective of the different families and genera (Flower, 1950; Sweet, 1964). The siphuncle in the Oncocerida is commonly located at or near the ventral margin. Connecting rings are mo ...
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Oncocerida
The Oncocerida comprise a diverse group of generally small nautiloid cephalopods known from the Middle Ordovician to the Mississippian (early Carboniferous; one possible member is known from the Early Permian), in which the connecting rings are thin and siphuncle segments are variably expanded (Flower, 1950). At present the order consists of some 16 families, a few of which, such as the Oncoceratidae, Brevicoceratidae, and Acleistoceratidae contain a fair number of genera each while others like the Trimeroceratidae and Archiacoceratidae are represented by only two or three (Sweet, 1964). Physical characteristics The shells of oncocerids are primarily somewhat compressed cyrtoconic brevicones. More advanced forms include gyrocones, serpenticones, torticones, and elongate orthocones and cyrtocones, reflective of the different families and genera (Flower, 1950; Sweet, 1964). The siphuncle in the Oncocerida is commonly located at or near the ventral margin. Connecting rings are mo ...
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Nautiloids
Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and ''Allonautilus''. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms ( orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day. In a broad sense, "nautiloid" refers to a major cephalopod subclass or collection of subclasses (Nautiloidea ''sensu lato''). Nautiloids are typically considered one of three main groups of cephalopods, along with the extinct ammonoids (ammonites) and living coleoids (such as squid, octopus, and kin). While ammonoids and coleoids are monophyletic clades with exclusive ancestor-descendant rela ...
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Graciloceratidae
The Graciloceratidae is a family of nautiloid cephalopods from the Middle and Upper Ordovician belonging to the Oncocerida, characterized by exogastric cyrtocones that expand slightly or moderately and have thin walled, orthochoanitic marginal or subventral, tubular siphuncles (Sweet 1964). The Graciloceratidae was proposed by Rousseau Flower in 1950 and at that time placed in the Basslerocerida which has since been abandoned. It is now assigned to the Oncocerida (Sweet 1964). Phylogeny The Graciloceratidae are most likely derived from the Bassleroceratidae (Flower 1950) from which they differ primarily in having thin rather than thick connecting rings. They are also the most likely source for the Oncoceratidae, Tripteroceratidae, and Valcouroceratidae (Flower 1950). Included genera The Graciloceratidae contains five known genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the h ...
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Valcouroceratidae
The Valcouroceratidae is a family within the Oncocerida, nautiloid cephalopods from the middle and upper Ordovician, established by Rousseau Flower in 1945. Diagnosis Valcouroceratids are characterized by exogastric cyrtocones and brevicones that change during the life span from compressed to depressed or subtriangular in cross section, and which have ventral siphuncles that are cyrtochoanitic and which contain lamellar actinosiphonate fillings. The actinosiphonate fillings, commonly referred to as deposits, are radially inward projections of the connecting rings that extend longitudinally along the inner wall of the siphuncle and project forward as blades into the siphuncle interior from the septal foramina.(Sweet 1964, Flower 1950) Distribution Most valcouroceratids come from North America but Valcouroceras has been found in northern Europe (Norway) as well. (Sweet 1964) Genera The Valcouroceratidae includes 8 genera: *'' Valcouroceras'' *'' Actinomorpha'' *'' Augustoceras'' ...
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Polyelasmoceratidae
Polyelasmoceratidae is a family of oncocerid nautiloids characterized by rapidly expanding endogastrically curved shells, curved such that the ventral side is longitudinally concave. In cross section shells are typically tear-drop in shape to subtriangular. The siphuncle is typically nummuloidal, like a string of beads, with outwardly flared septal necks, and located between the center and the venter. Most contain radially lamellar actinosiphonate deposits. The Middle Silurian ''Danaoceras ''Danaoceras'' is a nautiloid cephalopod from the middle Silurian of central Europe included in the oncoceroid family Polyelasmoceratidae. Similar specimens from the middle Devonian of North America may belong. ''Danaoceras'' has a laterally co ...'' is probably the ancestral genus, which give rise to the contemporary '' Codoceras'' and ultimately the subsequent Devonian genera. References * Walter C. Sweet, 1964. Nautiloidea - Oncocerida; Treatise on Invertebrate Paleontology, Part K. G ...
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Oncoceratidae
Oncoceratidae is a family of nauatiloid cephalopods in the order Oncocerida established by Hyatt, 1884, that range from the Middle Ordovician to the Upper Silurian. Diagnosis Oncoceratidae are characterized by generally compressed, cyrtoconic, and breviconic shells with an exogastric curvature such that the ventral profile is convex or more so than dorsal, and in which the siphuncle is generally empty and located ventral of the center. In primitive forms, the siphuncle in early growth stages is composed of tubular segments with almost straight suborthochoanitic septal necks, but becomes cyrtochoanitic with expanded segments in the later growth stages, and is expanded and cyrtochoanitic throughout in advanced forms. In a few advanced forms, the siphuncle is actinosiphonate. (Flower 1950, Sweet 1964) Evolution The Oncoceratidae first appeared early in the Middle Ordovician (Sweet 1964) simultaneously with the Graciloceratidae, Tripteroceratidae, and Valcouroceratidae, deri ...
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Karoceratidae
Karoceratidae is a family of nautiloids within the order Oncocerida The Oncocerida comprise a diverse group of generally small nautiloid cephalopods known from the Middle Ordovician to the Mississippian (early Carboniferous; one possible member is known from the Early Permian), in which the connecting rings are t ..., characterized by straight or curved, laterally narrow, shells and slender, ventral siphuncles that are empty except in '' Karoceras''. Siphuncle segments are inflated ventrally but straight dorsally. Septal necks are cyrtochoanitic, outwardly curved; connecting rings are thin. Genera include '' Karoceras'', '' Osbornoceras'', and possibly '' Shuranoceras'', which have been found in sediments ranging from the Lower Silurian to possibly as young as the Lower Devonian. References * Walter C. Sweet, 1964. Nautiloidea - Oncocerida; Treatise on Invertebrate Paleontology, Part K. Geological Society of America and University of Kansas Press. Prehistoric nautiloid f ...
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Tripteroceratidae
The Tripteroceratidae is a family of depressed, straight to slightly curved nautiloid cephalopods from the middle and upper Ordovician with generally flattened venters and empty siphuncles with straight to inflated segments included in the Oncocerida (Sweet, 1964). The Tripteroceratidae appeared almost simultaneously with the Oncoceratidae and Valcouroceratidae early in the Middle Ordovician, and are most likely derived from the Graciloceratidae The Graciloceratidae is a family of nautiloid cephalopods from the Middle and Upper Ordovician belonging to the Oncocerida, characterized by exogastric cyrtocones that expand slightly or moderately and have thin walled, orthochoanitic marginal .... Genera The Tripteroceratdae includes five known genera, as briefly described. ''Tripteroceras'', the type genus, named by Hyatt (1884) is characterized by a small, straight or slightly exogastric shell with a broad, depressed triangular cross section. The venter, underneath, is flat ...
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Hemiphragmoceratidae
Hemiphragmoceratidae is a family of endogastrically brevconic oncocerids characterized by elaborately visored apertures in which the hyponomic sinus in mature specimens is on a spout-like process and there may be lateral and dorsal salients. (Sweet 1964, Flower 1950). Shells are compressed with the apical portion curved and the anterior straight. Siphucles are nummuloideal with expanded spheroidal segments and continuously actinosiphonate interiors. The Hemiphragmoceratidae are probably descended from the Oncoceratidae and are known from the middle and upper Silurian but may range into the middle Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ... (Sweet 1964). They are similar with regards to their constricted and ornate apertures to the middle Silurian exogastric Trime ...
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Poterioceratidae
Poterioceratidae is a family of nautiloid cephalopods included in the Oncocerida that lived during the period from the Early Devonian to the Early Carboniferous ( Mississippian). Members of the Poterioceratidae are distinguished by a subcircular to compressed exogastric shell (curved upwardly so as the lower side, the venter, is convex viewed from the side) that has no hyponomic sinus and a central to subcentral siphuncle composed of subquadrate to nummuloidal segments in which the septal necks are more strongly curved on the upper, or dorsal side. This is opposite from the Karoceratidae in which siphuncle segments are inflated ventrally but straight dorsally. Some poterioceratid genera have actinosiphonate structures or annular deposits within the siphuncle. In others it is empty. The origin of the Poterioceratidae is undetermined but may have their origin in some Silurian member of the Acleistoceratidae or perhaps even in some Ordovician oncoceratid through an unknown Siluri ...
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Nothoceratidae
Nothoceratidae is a family of nautiloid cephalopods in the orthoceratoid order Oncocerida in which shells are exogastrically or endogastrically breviconic, planospiral, or torticonic; often with a constricted or visored aperture; and a siphuncle commonly composed of concave segments and occupied by actinosiphonate deposits. Some ten genera have been described which lived during the time between the Early Silurian and Late Devonian. The ancestral form is probably '' Perimecoceras'' which is known from the Upper Silurian of central Europe and which is similar in external form the ''Oonoceras'' from the Oncoceratidae. Genera *'' Blakeoceras'' *'' Bolloceras'' *'' Conostichoceras'' *'' Cyrthoceratites'' *'' Lorieroceras'' *'' Metaphragmoceras'' *'' Mutoblakeoceras'' *'' Nothoceras'' *'' Paraconradoceras'' *'' Perimecoceras'' *'' Tafilaltoceras'' *'' Turnoceras'' References * Sweet, W. C. 1964; Nautiloidea -Oncocerida; Treatise on Invertebrate Paleontology The ''Treatise ...
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Nautilida
The Nautilida constitute a large and diverse order of generally coiled nautiloid cephalopods that began in the mid Paleozoic and continues to the present with a single family, the Nautilidae which includes two genera, ''Nautilus'' and ''Allonautilus'', with six species. All told, between 22 and 34 families and 165 to 184 genera have been recognised, making this the largest order of the subclass Nautiloidea. Classification and phylogeny Current classification The current classification of the Nautilida, in prevalent use, is that of Bernhard Kummel (Kummel 1964) in the Treatise which divides the Nautilida into five superfamilies, the Aipocerataceae, Clydonautilaceae, Tainocerataceae, and Trigonocerataceae, mostly of the Paleozoic, and the later Nautilaceae. These include 22 families and some 165 or so genera (Teichert and Moore 1964) Other concepts Shimansky 1962 (in Kummel 1964) divided the Nautilida into five suborders, the mostly Paleozoic Centroceratina, Liroceratina, Rutoc ...
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