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Nautilida
The Nautilida constitute a large and diverse order of generally coiled nautiloid cephalopods that began in the mid Paleozoic and continues to the present with a single family, the Nautilidae which includes two genera, ''Nautilus'' and ''Allonautilus'', with six species. All told, between 22 and 34 families and 165 to 184 genera have been recognised, making this the largest order of the subclass Nautiloidea. Classification and phylogeny Current classification The current classification of the Nautilida, in prevalent use, is that of Bernhard Kummel (Kummel 1964) in the Treatise which divides the Nautilida into five superfamilies, the Aipocerataceae, Clydonautilaceae, Tainocerataceae, and Trigonocerataceae, mostly of the Paleozoic, and the later Nautilaceae. These include 22 families and some 165 or so genera (Teichert and Moore 1964) Other concepts Shimansky 1962 (in Kummel 1964) divided the Nautilida into five suborders, the mostly Paleozoic Centroceratina, Liroceratina, Rutoc ...
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Nautilidae
The nautilus (, ) is a pelagic marine mollusc of the cephalopod family Nautilidae. The nautilus is the sole extant family of the superfamily Nautilaceae and of its smaller but near equal suborder, Nautilina. It comprises six living species in two genera, the type genus, type of which is the genus ''Nautilus (genus), Nautilus''. Though it more specifically refers to species ''chambered nautilus, Nautilus pompilius'', the name chambered nautilus is also used for any of the Nautilidae. All are protected under CITES Appendix II. Depending on species, adult shell diameter is between 4 and 10 inches. Nautilidae, both extant and extinct, are characterized by involute or more or less convolute shells that are generally smooth, with compressed or depressed whorl (mollusc), whorl sections, straight to sinuous sutures, and a tubular, generally central siphuncle.Kümmel, B. 1964. Nautiloidae-Nautilida, in the Treatise on Invertebrate Paleontology, Geological Society of America and Univ of K ...
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Nautilus
The nautilus (, ) is a pelagic marine mollusc of the cephalopod family Nautilidae. The nautilus is the sole extant family of the superfamily Nautilaceae and of its smaller but near equal suborder, Nautilina. It comprises six living species in two genera, the type of which is the genus ''Nautilus''. Though it more specifically refers to species ''Nautilus pompilius'', the name chambered nautilus is also used for any of the Nautilidae. All are protected under CITES Appendix II. Depending on species, adult shell diameter is between 4 and 10 inches. Nautilidae, both extant and extinct, are characterized by involute or more or less convolute shells that are generally smooth, with compressed or depressed whorl sections, straight to sinuous sutures, and a tubular, generally central siphuncle.Kümmel, B. 1964. Nautiloidae-Nautilida, in the Treatise on Invertebrate Paleontology, Geological Society of America and Univ of Kansas Press, Teichert and Moore eds. Having survived relatively u ...
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Nautilus Subtruncatus
The nautilus (, ) is a pelagic marine mollusc of the cephalopod family Nautilidae. The nautilus is the sole extant family of the superfamily Nautilaceae and of its smaller but near equal suborder, Nautilina. It comprises six living species in two genera, the type of which is the genus ''Nautilus''. Though it more specifically refers to species ''Nautilus pompilius'', the name chambered nautilus is also used for any of the Nautilidae. All are protected under CITES Appendix II. Depending on species, adult shell diameter is between 4 and 10 inches. Nautilidae, both extant and extinct, are characterized by involute or more or less convolute shells that are generally smooth, with compressed or depressed whorl sections, straight to sinuous sutures, and a tubular, generally central siphuncle.Kümmel, B. 1964. Nautiloidae-Nautilida, in the Treatise on Invertebrate Paleontology, Geological Society of America and Univ of Kansas Press, Teichert and Moore eds. Having survived relatively un ...
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Nautiloidea
Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and ''Allonautilus''. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms ( orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day. In a broad sense, "nautiloid" refers to a major cephalopod subclass or collection of subclasses (Nautiloidea ''sensu lato''). Nautiloids are typically considered one of three main groups of cephalopods, along with the extinct ammonoids (ammonites) and living coleoids (such as squid, octopus, and kin). While ammonoids and coleoids are monophyletic clades with exclusive ancestor-descendant rela ...
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Nautilina
The Nautilina is the last suborder of the Nautilida and the only nautiloids living since the end of the Triassic. The Nautilina, proposed by Shimanskiy, is basically the Nautilaceae of Kummel, 1964, defined by Furnish and Glenister, but differs in omitting two families, the Paracenoceratidae and Pseudonautilidae which instead are placed in the Liroceratina. The Nautilina are derived from the Syringonautilidae, a family in the Centroceratina (Trigonocerataceae), in the Late Triassic and consists of four families, the Nautilidae, Cymatoceratidae, Herocoglossidae, and Aturiidae. The Nautilidae, which is the root stock of the suborder, includes the living ''Nautilus''. The Nautilidae are involute or slightly evolute and generally smooth with straight to sinuous sutures. The Cymatoceratidae, which are the most common of the Cretaceous nautiloids, are strongly ribbed. The Hercoglossidae are smooth but with differentiated sutures, in some with deep lateral lobes and well-developed ...
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Rutocerina
The Rutocertina is one of only three suborders in Shimankiy's (1957) classification of the Nautilida, the other two being the Lirocerina and Nautilina. Genera in the Rutocerina are redistributed (Kümmel 1964) in the Rutoceratina, Tainoceratina, and Centroceratina. The Lirocerina is redefined as the Liroceratina, and Nautilina.remains as is. In general terms these are similar to the simpler classification proposed by Kümmel 1964, wherein the Nautilida is divided into five superfamilies, the Tainocerataceae, Trigonocerataceae, Clydonautilacea, Aipocerataceae, and Nautilaceae. Shimanskiy's classification involves 34 families, Kümmel's only twenty-seven. The Rutoceratina (Shmanskiy 1957) is divided into two unequal superfamilies which do not correspond with the superfamilies of Kümmel, 1964. They are the Rutoceratacea and the Solenochilaceae. The Rutoceratacea is essentially the Rutoceratidae of Kümmel 1964, elevated and expanded to contain two families, the Rutoceratidae in ...
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Rutoceratidae
Rutoceratidae is a family of prototypical nautilids, derived probably from either Brevicoceratidae or Acleistoceratidae of the order Oncocerida early in the Devonian. Rutoceratidae comprise a family within the oncocerid superfamily TainocerataceaeKummel 1964, Nautiloidea-Nautilida; Treatise on Invertebrate Paleontology, Part K: Nautiloidea, Teichert & Moore (eds) They are generally characterized by cyrtoconic and gyroconic shells, commonly with spines, nodes, or frills, although some included genera are almost orthoconic, and a commonly empty, tubular ventral siphuncle. The Rutocertids lived during the Devonian and Mississippian (early Carboniferous) and are the ancestral stock of Nautilida. Within the superfamily Taintocerataceae, rutoceratids gave rise to the exclusively Devonian family Tetragonoceratidae and near the start of the Mississippian to the family Koninckioceratidae which lasted into the Permian and to Tainoceratidae which lasted through most of the Triassic. ...
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Centroceratina
The Trigonoceratoidea are a superfamily within the Nautilida that ranged from the Devonian to the Triassic, thought to have contained the source for the Nautilaceae in which ''Nautilus'' is found. Trigonoceratoidea are characterized by open-spiraled, gyroconic, to closed, nautiliconic shells in which the Whorl (mollusc), Whorl section is wikt:quadrate#Adjective, quadrate in primitive forms; the wikt:venter#Etymology 2, venter typically narrow to acute, the dorsum broad. In some advanced forms, the venter may become concave or broad and rounded, and in some, the surfaces may be strongly lirate. Classification and phylogeny The Trigonoceratoidea are based on the family Trigonoceratidae of Alpheus Hyatt, 1884, with which other phylogenetically related families are combined, and are equivalent to the abandoned Centroceratida of Flower in Flower and Kümmel 1950, and to the Centroceratina of Shimanskiy 1957, revised to the Centrocerataceae, Shimanskiy 1962. The Trigonoceratoidea comb ...
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Tainoceratina
''For a further discussion of this group as in use today, see the superfamily Tainocerataceae'' The Tainoceratina is a suborder within the Nautilida (Kümmel 1964) created by Shimanskiy on the assumption that the initial, or embryonic, whorl was the critical phylogenetic indicator, which resulted in combining otherwise unrelated and separating obviously related forms. The Tainoceratina rearranges the precedented Tainocerataceae The Tainoceratoidea are a superfamily in the cephalopod order Nautilida characterized by straight to loosely coiled shells, generally to a degree such that the width is greater than the height, to quadrate whorl section. Many bore spines, ribs, f ... while excluding the ancestral Rutoceratidae. Three families, the Tainoceratidae, Tetragonoceratidae, and Rhiphaeoceratidae are in common with the Tainocerataeae, but with somewhat different meaning and content. Six additional families used in the Tainoceratina, the Actubonautilidae, Encoiloceratidae, Gzhel ...
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Oncocerida
The Oncocerida comprise a diverse group of generally small nautiloid cephalopods known from the Middle Ordovician to the Mississippian (early Carboniferous; one possible member is known from the Early Permian), in which the connecting rings are thin and siphuncle segments are variably expanded (Flower, 1950). At present the order consists of some 16 families, a few of which, such as the Oncoceratidae, Brevicoceratidae, and Acleistoceratidae contain a fair number of genera each while others like the Trimeroceratidae and Archiacoceratidae are represented by only two or three (Sweet, 1964). Physical characteristics The shells of oncocerids are primarily somewhat compressed cyrtoconic brevicones. More advanced forms include gyrocones, serpenticones, torticones, and elongate orthocones and cyrtocones, reflective of the different families and genera (Flower, 1950; Sweet, 1964). The siphuncle in the Oncocerida is commonly located at or near the ventral margin. Connecting rings are mo ...
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Allonautilus
The genus ''Allonautilus'' contains two species of nautiluses, which have a significantly different morphology from those placed in the sister taxon ''Nautilus''. ''Allonautilus'' is now thought to be a descendant of ''Nautilus'', rendering the latter genus paraphyletic. Live individuals of the genus have only been collected in Papua New Guinea and the Solomon Islands. Little is known about their biology because they live in deep waters, whereas the better-understood genus ''Nautilus'' lives closer to the surface. The entire family Nautilidae, including all species in the genus ''Nautilus'' and ''Allonautilus'', was listed on Appendix II of the Convention on International Trade in Endangered Species of Wild Fauna and Flora CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of intern ... (CITE ...
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Centroceratidae
The Centroceratidae is the ancestral family of the Trigonoceratoidea and of the equivalent Centroceratina; extinct shelled cephalopods belonging to the order Nautilida Diagnosis The Centroceratidae, which range from the Middle Devonian to the Lower Permian, are characterized by gyroconic, evolute tarphyceraconic, and involute nautiliconic shells with compressed whorls, typically with a quadrangular whorl section in which the flanks converge on venter that is much narrower than the dorsum and ventral and umbilical shoulders are sharply angular, or rarely rounded. In some, e.g. ''Centroceras'', the flanks are divided by a ridge that runs along the middle. Sutures have ventral and lateral lobes but are transverse dorsally. The siphuncle is tubular and close to but not on in contact with the venter (Kummel 1964). Phylogeny and genera Evolutionary sequence The Centroceratidae are thought probably to be derived from the rutoceratid stock (Flower 1950, 1988, Kummel 1964) Earliest are ...
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