Guadalupian Synapsids
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Guadalupian Synapsids
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lop ...
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International Commission On Stratigraphy
The International Commission on Stratigraphy (ICS), sometimes referred to unofficially as the "International Stratigraphic Commission", is a daughter or major subcommittee grade scientific daughter organization that concerns itself with stratigraphy, stratigraphical, geology, geological, and chronology, geochronological matters on a global scale. It is the largest subordinate body of the International Union of Geological Sciences (IUGS). The ICS is essentially a permanent working committee, working subcommittee, which meets far more regularly than the quadrennial meetings scheduled by the IUGS, when it meets as a congress or committee, membership of the whole. Aims One of its main aims, a project begun in 1974, is to establish a multidisciplinary standard and global geologic time scale that will ease paleontology, paleontological and geobiology, geobiological comparisons region to region by benchmarks with stringent and rigorous strata criteria called Global Boundary Stratotype ...
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Lopingian
The Lopingian is the uppermost series/last epoch of the Permian. It is the last epoch of the Paleozoic. The Lopingian was preceded by the Guadalupian and followed by the Early Triassic. The Lopingian is often synonymous with the informal terms late Permian or upper Permian. The name was introduced by Amadeus William Grabau in 1931 and derives from Leping, Jiangxi in China. It consists of two stages/ ages. The earlier is the Wuchiapingian and the later is the Changhsingian. The International Chronostratigraphic Chart (v2018/07) provides a numerical age of 259.1 ±0.5 Ma. If a Global Boundary Stratotype Section and Point (GSSP) has been approved, the lower boundary of the earliest stage determines numerical age of an epoch. The GSSP for the Wuchiapingian has a numerical age of 259.8 ± 0.4 Ma. Evidence from Milankovitch cycles suggests that the length of an Earth day during this epoch was approximately 22 hours. The Lopingian ended with the Permian–Triassic extinction event. ...
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Mass Extinction
An extinction event (also known as a mass extinction or biotic crisis) is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp change in the diversity and abundance of multicellular organisms. It occurs when the rate of extinction increases with respect to the background extinction rate and the rate of speciation. Estimates of the number of major mass extinctions in the last 540 million years range from as few as five to more than twenty. These differences stem from disagreement as to what constitutes a "major" extinction event, and the data chosen to measure past diversity. The "Big Five" mass extinctions In a landmark paper published in 1982, Jack Sepkoski and David M. Raup identified five particular geological intervals with excessive diversity loss. They were originally identified as outliers on a general trend of decreasing extinction rates during the Phanerozoic, but as more stringent statistical tests have been applied to ...
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Titanophoneus Chart2 DB15
''Titanophoneus'' ("titanic murderer") is an extinct genus of carnivorous dinocephalian therapsid from the Middle Permian. It is classified within the family Anteosauridae. The type species is ''Titanophoneus potens''. Remains of ''Titanophoneus'' have been found at Isheevo in Russia. Description An adult skull would have reached 80 cm with a heavy long snout. The long tail and short limbs show the species to be a primitive therapsid unlike ''Inostrancevia'' which was more advanced. The structure of the limbs and the density of the bone are designed for a sprawling stance. The temporal opening is more advanced than Estemmenosuchidae but less advanced than ''Inostrancevia''. The teeth are large with 12 large palate incisors followed by 2 canines and various smaller back teeth. The lower palate is the same as the upper but without the canine teeth. The appearance of ''Titanophoneus'' is reminiscent to the sphenacodontid pelycosaurs, which included '' Dimetrodon''. Gallery i ...
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Theriodont
The theriodonts (clade Theriodontia) are a major group of therapsids which appeared during the Middle Permian and which includes the gorgonopsians and the eutheriodonts, itself including the therocephalians and the cynodonts. Naming In 1876, Richard Owen named a suborder Theriodontia, which he divided into the Cynodontia and the Gomphodontia. The modern clade concept was devised by James Allen Hopson. In his system, Theriodontia fall into two main groups: the Gorgonopsia and the Eutheriodontia. The latter consist of the Therocephalia and Cynodontia.J.A. Hopson. 1999. "Therapsids". ''Encyclopedia of Paleontology'' 2: 1256-1266 Evolution Theriodonts appeared at the same time as their sister group within the Neotherapsida, the Anomodontia, about 270 million years ago, in the Middle Permian. Even these early theriodonts were more mammal-like than their anomodont and dinocephalian contemporaries. Early theriodonts may have been warm-blooded. Early forms were carnivorous, but se ...
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Biarmosuchia
Biarmosuchians are an extinct clade of non-mammalian synapsids from the Permian. They are the most basal group of the therapsids. All of them were moderately-sized, lightly-built carnivores, intermediate in form between basal sphenacodont "pelycosaurs" and more advanced therapsids. Biarmosuchians were rare components of Permian ecosystems, and the majority of species belong to the clade Burnetiamorpha, which are characterized by elaborate cranial ornamentation. Characteristics The biarmosuchian skull is very similar to the sphenacodontid skull, differing only in the larger temporal fenestra (although these are still small relative to later therapsids), slightly backward-sloping occiput (the reverse of the pelycosaur condition), reduced number of teeth, and single large canine teeth in both upper and lower jaws, and other features (Carroll 1988 pp. 370, Benton 2000 p. 114). In later specialised Biarmosuchia, these resemble the enlarged canines of the Gorgonopsia. The pr ...
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Anomodont
Anomodontia is an extinct group of non-mammalian therapsids from the Permian and Triassic periods. By far the most speciose group are the dicynodonts, a clade of beaked, tusked herbivores.Chinsamy-Turan, A. (2011) ''Forerunners of Mammals: Radiation - Histology - Biology''p.39.Indiana University Press, . Retrieved May 2012 Anomodonts were very diverse during the Middle Permian, including primitive forms like ''Anomocephalus'' and ''Patranomodon'' and groups like Venyukovioidea and Dromasauria. Dicynodonts became the most successful and abundant of all herbivores in the Late Permian, filling ecological niches ranging from large browsers down to small burrowers. Few dicynodont families survived the Permian–Triassic extinction event, but one lineage (Kannemeyeriiformes) evolved into large, stocky forms that became dominant terrestrial herbivores right until the Late Triassic, when changing conditions caused them to decline, finally going extinct during the Triassic-Jurassic ext ...
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Dinocephalia
Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included herbivorous, carnivorous, and omnivorous forms. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania. Description Apart from the Biarmosuchians, the dinocephalians are the least advanced therapsids, although still uniquely specialised in their own way. They retain a number of primitive characteristics (e.g. no secondary palate, small dentary) shared with their pelycosaur ancestors, although they are also more advanced in possessing therapsid adaptations like the expansion of the ilium and more erect limbs. They include carni ...
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Clades
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms t ...
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Sphenacodontia
Sphenacodontia is a stem-based clade of derived synapsids. It was defined by Amson and Laurin (2011) as "the largest clade that includes ''Haptodus baylei'', ''Haptodus garnettensis'' and '' Sphenacodon ferox'', but not ''Edaphosaurus pogonias''". They first appear during the Late Pennsylvanian (Upper Carboniferous) epoch. From the end of the Carboniferous to the end of the Permian, most of them remained large, with only some secondarily becoming small in size. Basal Sphenacodontia constitute a transitional evolutionary series from early pelycosaurs to ancestral therapsids (which in turn were the ancestors of more advanced forms and finally the mammals). One might say that the sphenacodontians are proto-therapsids (even though there is almost a 30-million-years gap between the separation of the ancestors of therapsids from other sphenacodontians and the first appearance of therapsids in the fossil record). Characteristics The defining characteristics include a thickening of ...
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Pelycosaur
Pelycosaur ( ) is an older term for basal or primitive Late Paleozoic synapsids, excluding the therapsids and their descendants. Previously, the term ''mammal-like reptile'' had been used, and pelycosaur was considered an order, but this is now thought to be incorrect, and seen as outdated. Because it excludes the advanced synapsid group Therapsida, the term is paraphyletic and contrary to modern formal naming practice. Thus the name ''pelycosaurs'', similar to the term ''mammal-like reptiles'', had fallen out of favor among scientists by the 21st century, and is only used informally, if at all, in the modern scientific literature. The terms stem mammals, protomammals, and basal or primitive synapsids are used where needed, instead. Etymology The term ''pelycosaur'' has been fairly well abandoned by paleontologists because it no longer matches the features that distinguish a clade. The modern word was created from Greek meaning 'wooden bowl' or 'axe' and meaning 'lizar ...
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Therapsid
Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more underneath the body, as opposed to the sprawling posture of many reptiles and salamanders. Therapsids evolved from "pelycosaurs", specifically within the Sphenacodontia, more than 279.5 million years ago. They replaced the "pelycosaurs" as the dominant large land animals in the Middle Permian through to the Early Triassic. In the aftermath of the Permian–Triassic extinction event, therapsids declined in relative importance to the rapidly diversifying reptiles during the Middle Triassic. The therapsids include the cynodonts, the group that gave rise to mammals ( Mammaliaformes) in the Late Triassic, around 225 million years ago. Of the non-mammalian therapsids, only cynodonts survived beyond the end of the Triassic, with the only other ...
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