The Guadalupian is the second and middle

series Series may refer to: People with the name * Caroline Series (born 1951), English mathematician, daughter of George Series * George Series (1920–1995), English physicist Arts, entertainment, and media Music * Series, the ordered sets used in ...

epoch In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided by ...

of the

Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...

. The Guadalupian was preceded by the

Cisuralian The Cisuralian is the first Series (stratigraphy), series/Epoch (geology), epoch of the Permian. The Cisuralian was preceded by the Pennsylvanian (geology), Pennsylvanian and followed by the Guadalupian. The Cisuralian Epoch is named after the w ...

and followed by the

Lopingian The Lopingian is the uppermost series/last epoch of the Permian. It is the last epoch of the Paleozoic. The Lopingian was preceded by the Guadalupian and followed by the Early Triassic. The Lopingian is often synonymous with the informal terms l ...

. It is named after the

Guadalupe Mountains The Guadalupe Mountains ( es, Sierra de Guadalupe) are a mountain range located in West Texas and southeastern New Mexico. The range includes the highest summit in Texas, Guadalupe Peak, , and the "signature peak" of West Texas, El Capitan, both ...

of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the

therapsid Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more ...

s, a minor extinction event called

Olson's Extinction Olson's Extinction was a mass extinction that occurred in the late Cisuralian or early Guadalupian of the Permian period and which predated the Permian–Triassic extinction event. It is named after Everett C. Olson. There was a sudden change be ...

and a significant mass extinction called the

end-Capitanian extinction event The Capitanian mass extinction event, also known as the end-Guadalupian extinction event or the pre-Lopingian crisis was an extinction event that predated the end-Permian extinction event and occurred around 260 million years ago during a period ...

. The Guadalupian was previously known as the Middle Permian.

Name and background

The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains in New Mexico. The International Chronostratigraphic Chart V2021/07 provides a numerical age of 273.01 ± 0.14 – 259.51 ± 0.21 mya.

Biodiversity

Therapsid Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more ...

s became the dominant land animals in Guadalupian, displacing the

pelycosaur Pelycosaur ( ) is an older term for basal or primitive Late Paleozoic synapsids, excluding the therapsids and their descendants. Previously, the term ''mammal-like reptile'' had been used, and pelycosaur was considered an order, but this is no ...

s. Therapsids evolved from a group of pelycosaurs called sphenacodonts. Therapsida consists of four major

clades A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, t ...

: the

dinocephalia Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinoceph ...

ns, the herbivorous

anomodont Anomodontia is an extinct group of non-mammalian therapsids from the Permian and Triassic periods. By far the most speciose group are the dicynodonts, a clade of beaked, tusked herbivores.Chinsamy-Turan, A. (2011) ''Forerunners of Mammals: Ra ...

s, the carnivorous

biarmosuchia Biarmosuchians are an extinct clade of non-mammalian synapsids from the Permian. They are the most basal group of the therapsids. All of them were moderately-sized, lightly-built carnivores, intermediate in form between basal sphenacodont "pelyc ...

ns, and the mostly carnivorous

theriodont The theriodonts (clade Theriodontia) are a major group of therapsids which appeared during the Middle Permian and which includes the gorgonopsians and the eutheriodonts, itself including the therocephalians and the cynodonts. Naming In 1876, Rich ...

s. After a brief burst of evolutionary diversity, the dinocephalians died out in the later Guadalupian. Titanophoneus chart2 DB15

mass extinction An extinction event (also known as a mass extinction or biotic crisis) is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp change in the diversity and abundance of multicellular organisms. It ...

occurred in the early Guadalupian before the larger

Permian–Triassic extinction event The Permian–Triassic (P–T, P–Tr) extinction event, also known as the Latest Permian extinction event, the End-Permian Extinction and colloquially as the Great Dying, formed the boundary between the Permian and Triassic geologic periods, as ...

. This extinction was originally called Olson's Gap because it was thought to be a problem in preservation of fossils. Since the 1990s it has been renamed Olson’s Extinction. This extinction event occurred near the beginning of the epoch and led to an extended period of low diversity when two-thirds of terrestrial vertebrate life was lost worldwide. Global diversity rose dramatically by the end probably the result of disaster taxa filling empty guilds, only to fall again when the end-Guadalupian event caused a diversity drop in the

Wuchiapingian In the geologic timescale, the Wuchiapingian or Wujiapingian (from in the Liangshan area of Hanzhong, Shaanxi Province) is an age or stage of the Permian. It is also the lower or earlier of two subdivisions of the Lopingian Epoch or Series. The ...

. There is no agreed cause for the Olson's Extinction.

Climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to E ...

may be a possible cause. Extreme environments were observed from the Permian of Kansas which resulted from a combination of hot climate and acidic waters particularly coincident with Olson’s Extinction. Whether this climate change was a result of Earth's natural processes or exacerbated by another event is unknown.

Climate

The climate resembled that of much of central Asia today.

Pangea Pangaea or Pangea () was a supercontinent that existed during the late Paleozoic and early Mesozoic eras. It assembled from the earlier continental units of Gondwana, Euramerica and Siberia during the Carboniferous approximately 335 million y ...

was a supercontinent and had very hot dry summers and cold bitter winters. The coasts were tropical and had monsoons. The first two-thirds of the epoch were the continuation of a temperate and tropical climate. This started to dry out and the coal forming of the previous epoch stopped. The change in climate also provided a new environment for new tetrapods, reptiles, fish, plants, and invertebrates. In the last third the temperature started to drop and many coral reefs died out. If that was not enough, increased volcano activity brought a reduction in oxygen, a greenhouse and mass extinction.

Beginning or Roadian

The

Roadian In the geologic timescale, the Roadian is an age or stage of the Permian. It is the earliest or lower of three subdivisions of the Guadalupian Epoch or Series. The Roadian lasted between and million years ago (Ma). It was preceded by the Kung ...

Stage was between 272.3 ± 0.5 – 268.8 ± 0.5 Mya.

Olson’s Extinction Olson's Extinction was a mass extinction that occurred in the late Cisuralian or early Guadalupian of the Permian period and which predated the Permian–Triassic extinction event. It is named after Everett C. Olson. There was a sudden change bet ...

was a worldwide loss of terrestrial vertebrate life that occurred during the Roadian and Wordian. Fauna did not recover fully from Olson's Extinction before the impact of the Permian-Triassic extinction event. Estimates of recovery time vary, where some authors indicated recovery was prolonged, lasting 30 million years into the Triassic. Several important events took place during Olson's Extinction, most notably the origin of

therapsids Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented mor ...

, a group that includes the evolutionary ancestors of mammals. Further research on the recently identified primitive therapsid of the

Xidagou Formation The Qingtoushan Formation is a Middle Permian-age geologic formation in the Qilian Mountains of Gansu, China. It is known for its diverse tetrapod fauna known as the Dashankou fauna, which likely dates to the Roadian, and includes some of the ol ...

(Dashankou locality) in China of Roadian age may provide more information on this topic.

Middle or Wordian

The

Wordian In the geologic timescale, the Wordian is an age or stage of the Permian. It is the middle of three subdivisions of the Guadalupian Epoch or Series. The Wordian lasted between and million years ago (Ma). It was preceded by the Roadian and foll ...

Stage was between 268.8 ± 0.5 – 265.1 ± 0.4 Mya. The base of the Wordian Stage is defined as the place in the stratigraphic record where fossils of

conodont Conodonts (Greek ''kōnos'', "cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which ...

species ''

Jinogondolella ''Jinogondolella'' is an extinct genus of conodonts.Mesogondolella and Jinogondolella (Conodonta): Multielement definition of the taxa that bracket the basal Guadalupian (Middle Permian Series) GSSP. Lance L. Lambert, Bruce R. Wardlaw and Charle ...

aserrata'' first appear. The global reference profile for this stratigraphic boundary is located at Getaway Ledge in the

Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...

. The top of the Wordian (the base of the Capitanian Stage) is defined as the place in the stratigraphic record where the conodont species ''Jinogondolella postserrata'' first appears.

Conclusion or Capitanian

The

Capitanian In the geologic timescale, the Capitanian is an age or stage of the Permian. It is also the uppermost or latest of three subdivisions of the Guadalupian Epoch or Series. The Capitanian lasted between and million years ago. It was preceded by th ...

Stage was between 265.1 ± 0.4 – 259.8 ± 0.4 Mya. The Guadalupian ended with a deteriorating environment, Greenhouse conditions, and several series of mass-extinctions; both the great

dinocephalian Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocepha ...

s, other taxa on land and various invertebrates in the sea. They would be succeeded by new types of therapsids, especially the

gorgonopsia Gorgonopsia (from the Greek Gorgon, a mythological beast, and 'aspect') is an extinct clade of sabre-toothed therapsids from the Middle to Upper Permian roughly 265 to 252 million years ago. They are characterised by a long and narrow skull, a ...

ns among others. A significant mass extinction event (the

End-Capitanian extinction event The Capitanian mass extinction event, also known as the end-Guadalupian extinction event or the pre-Lopingian crisis was an extinction event that predated the end-Permian extinction event and occurred around 260 million years ago during a period ...

) occurred at the end of this epoch, which was associated with anoxia and

acidification Acidification may refer to: * Ocean acidification, decrease in the pH of the Earth's oceans * Freshwater acidification, atmospheric depositions and soil leaching of SOx and NOx * Soil acidification, buildup of hydrogen cations, which reduces the ...

in the oceans and possibly caused by the volcanic eruptions that produced the

Emeishan Traps The Emeishan Traps constitute a flood basalt volcanic province, or large igneous province, in south-western China, centred in Sichuan province. It is sometimes referred to as the Permian Emeishan Large Igneous Province or Emeishan Flood Basalts. Li ...

. This extinction event may be related to the much larger

that followed about 10 million years later.

Carbon isotope Carbon (6C) has 15 known isotopes, from to , of which and are stable. The longest-lived radioisotope is , with a half-life of years. This is also the only carbon radioisotope found in nature—trace quantities are formed cosmogenically by t ...

s in

marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...

limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...

from the Capitanian Age show an increase in δ¹³C values. The change in carbon isotopes in the sea water reflects cooling of global climates. This climatic cooling may have caused the end-Capitanian extinction event among species that lived in warm water, like larger

fusulinid The Fusulinida is an extinct order within the Foraminifera in which the tests are traditionally considered to have been composed of microgranular calcite. Like all forams, they were single-celled organisms. In advanced forms the test wall was di ...

s (Verbeekninidae), large

bivalve Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bival ...

s (

Alatoconchidae Alatoconchidae is an extinct family of prehistoric bivalves that lived in the early to middle Permian period. Genera belonging to Alatoconchidae are characterized by their shell that is strongly compressed in the dorsoventral direction. Some spe ...

) and

rugose Rugose means "wrinkled". It may refer to: * Rugosa, an extinct order of coral, whose rugose shape earned it the name * Rugose, adjectival form of rugae Species with "rugose" in their names * '' Idiosoma nigrum'', more commonly, a black rugose tr ...

corals, and Waagenophyllidae.

Other subdivisions

Subdivisions that are sometimes used are, :*Kazanian or Maokovian (European) 70.6 ± 0.7 – 260.4 ± 0.7 Myaref> :*Braxtonian Stage (New Zealand) 70.6 ± 0.7 – 260.4 ± 0.7 Mya

References

{{Geological history of Earth, p, m Geological epochs *02