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Cyclocephala Castanea
''Cyclocephala castanea'' is a species of small beetle in the family Scarabaeidae which is native to the Amazon basin in South America. This beetle and certain others in the same genus have a commensal relationship with the large water lily '' Victoria amazonica''. Visiting the flower for food, the beetles are trapped inside and emerge laden with pollen the following evening. Ecology ''Cyclocephala castanea'', and the closely related '' Cyclocephala hardyi'', have co-evolved with the giant water lily '' Victoria amazonica'', found in shallow waters in the Amazon basin. The flowers are nocturnal and float on the surface of the water. They are creamy-white and have a strong fragrance when they first open. The beetles are attracted to the flowers which have a high temperature, up to 10 °C (18 °F) warmer than their surroundings, and which contain starch-rich appendages called paracarpels. At the end of the night the flowers close and the beetles are trapped by the pet ...
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Beetle
Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 described species, is the largest of all orders, constituting almost 40% of described insects and 25% of all known animal life-forms; new species are discovered frequently, with estimates suggesting that there are between 0.9 and 2.1 million total species. Found in almost every habitat except the sea and the polar regions, they interact with their ecosystems in several ways: beetles often feed on plants and fungi, break down animal and plant debris, and eat other invertebrates. Some species are serious agricultural pests, such as the Colorado potato beetle, while others such as Coccinellidae (ladybirds or ladybugs) eat aphids, scale insects, thrips, and other plant-sucking insects that damage crops. Beetles typically have a particularly hard e ...
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Scarabaeidae
The family Scarabaeidae, as currently defined, consists of over 30,000 species of beetles worldwide; they are often called scarabs or scarab beetles. The classification of this family has undergone significant change in recent years. Several subfamilies have been elevated to family rank (e.g., Bolboceratidae, Geotrupidae, Glaresidae, Glaphyridae, Hybosoridae, Ochodaeidae, and Pleocomidae), and some reduced to lower ranks. The subfamilies listed in this article are in accordance with those in Bouchard (2011). Description Scarabs are stout-bodied beetles, many with bright metallic colours, measuring between . They have distinctive, clubbed antennae composed of plates called lamellae that can be compressed into a ball or fanned out like leaves to sense odours. Many species are fossorial, with legs adapted for digging. In some groups males (and sometimes females) have prominent horns on the head and/or pronotum to fight over mates or resources. The largest fossil scaraba ...
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Amazon Basin
The Amazon basin is the part of South America drained by the Amazon River and its tributaries. The Amazon drainage basin covers an area of about , or about 35.5 percent of the South American continent. It is located in the countries of Bolivia, Brazil, Colombia, Ecuador, Guyana, Peru, Suriname, and Venezuela. Most of the basin is covered by the Amazon rainforest, also known as Amazonia. With a area of dense tropical forest, this is the largest rainforest in the world.   Geography The Amazon River begins in the Andes Mountains at the west of the basin with its main tributary the Marañón River and Apurimac River in Peru. The highest point in the watershed of the Amazon is the second biggest peak of Yerupajá at . With a length of about before it drains into the Atlantic Ocean, it is one of the two longest rivers in the world. A team of scientists has claimed that the Amazon is longer than the Nile, but debate about its exact length continues. The Amazon system ...
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South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southern subregion of a single continent called America. South America is bordered on the west by the Pacific Ocean and on the north and east by the Atlantic Ocean; North America and the Caribbean Sea lie to the northwest. The continent generally includes twelve sovereign states: Argentina, Bolivia, Brazil, Chile, Colombia, Ecuador, Guyana, Paraguay, Peru, Suriname, Uruguay, and Venezuela; two dependent territories: the Falkland Islands and South Georgia and the South Sandwich Islands; and one internal territory: French Guiana. In addition, the ABC islands of the Kingdom of the Netherlands, Ascension Island (dependency of Saint Helena, Ascension and Tristan da Cunha, a British Overseas Territory), Bouvet Island ( dependency of Norway), Pa ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Commensalism
Commensalism is a long-term biological interaction (symbiosis) in which members of one species gain benefits while those of the other species neither benefit nor are harmed. This is in contrast with mutualism, in which both organisms benefit from each other; amensalism, where one is harmed while the other is unaffected; parasitism, where one is harmed and the other benefits, and parasitoidism, which is similar to parasitism but the parasitoid has a free-living state and instead of just harming its host, it eventually ends up killing it. The commensal (the species that benefits from the association) may obtain nutrients, shelter, support, or locomotion from the host species, which is substantially unaffected. The commensal relation is often between a larger host and a smaller commensal; the host organism is unmodified, whereas the commensal species may show great structural adaptation consistent with its habits, as in the remoras that ride attached to sharks and other fishes. Remo ...
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Victoria Amazonica
''Victoria amazonica'' is a species of flowering plant, the second largest of the water lily family Nymphaeaceae. It is the national flower of Guyana. Its native regions are Guyana and tropical South America. Description The ''Victoria amazonica'' has very large leaves, up to in diameter, that float on the water's surface on a submerged stalk, in length, rivaling that of the green anaconda, a snake local to its habitat. It is the second-largest waterlily in the world. ''V. amazonica'' is native to the shallow waters of the Amazon River basin, such as oxbow lakes and bayous. In their native habitat, the flowers first begin to open as the sun starts to set and can take up to 48 hours to fully open. These flowers can grow up to 40 cm (16 in) in diameter. Each plant continues to produce flowers for a full growing season, and they have co-evolved a mutualistic relationship with a species of scarab beetle of the genus '' Cyclocephala'' as a pollinator. All the bud ...
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Victoria Regia
''Victoria amazonica'' is a species of flowering plant, the second largest of the water lily family Nymphaeaceae. It is the national flower of Guyana. Its native regions are Guyana and tropical South America. Description The ''Victoria amazonica'' has very large leaves, up to in diameter, that float on the water's surface on a submerged stalk, in length, rivaling that of the green anaconda, a snake local to its habitat. It is the second-largest waterlily in the world. ''V. amazonica'' is native to the shallow waters of the Amazon River basin, such as oxbow lakes and bayous. In their native habitat, the flowers first begin to open as the sun starts to set and can take up to 48 hours to fully open. These flowers can grow up to 40 cm (16 in) in diameter. Each plant continues to produce flowers for a full growing season, and they have co-evolved a mutualistic relationship with a species of scarab beetle of the genus '' Cyclocephala'' as a pollinator. All the buds ...
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Cyclocephala Hardyi
''Cyclocephala'' is a genus of scarab beetles from the subfamily Dynastinae (Coleoptera, Scarabaeidae). Beetles of this genus occur from southeastern Canada to Argentina and the West Indies. Adults of this genus are nocturnal or crepuscular, and are usually attracted to lights. Selected species This is a large genus and new species continue to be added. *'' Cyclocephala almitana'' *'' Cyclocephala amazona'' *''Cyclocephala amblyopsis'' *''Cyclocephala atripes'' *''Cyclocephala atripes'' *'' Cyclocephala barroensis'' *'' Cyclocephala brittoni'' *'' Cyclocephala borealis'' - Northern masked chafer *''Cyclocephala carbonaria'' *''Cyclocephala cartwrighti'' *'' Cyclocephala castanea'' *'' Cyclocephala castaniella'' *'' Cyclocephala colasi'' *'' Cyclocephala complanata'' *'' Cyclocephala concolor'' *'' Cyclocephala confusa'' *'' Cyclocephala conspicua'' *'' Cyclocephala discicollis'' *'' Cyclocephala discolor'' *'' Cyclocephala elegans'' *''Cyclocephala epistomalis'' *'' ...
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Coevolution
In biology, coevolution occurs when two or more species reciprocally affect each other's evolution through the process of natural selection. The term sometimes is used for two traits in the same species affecting each other's evolution, as well as gene-culture coevolution. Charles Darwin mentioned evolutionary interactions between flowering plants and insects in ''On the Origin of Species'' (1859). Although he did not use the word coevolution, he suggested how plants and insects could evolve through reciprocal evolutionary changes. Naturalists in the late 1800s studied other examples of how interactions among species could result in reciprocal evolutionary change. Beginning in the 1940s, plant pathologists developed breeding programs that were examples of human-induced coevolution. Development of new crop plant varieties that were resistant to some diseases favored rapid evolution in pathogen populations to overcome those plant defenses. That, in turn, required the development of ...
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Stigma (botany)
The stigma () is the receptive tip of a carpel, or of several fused carpels, in the gynoecium of a flower. Description The stigma, together with the style and ovary (typically called the stigma-style-ovary system) comprises the pistil, which is part of the gynoecium or female reproductive organ of a plant. The stigma itself forms the distal portion of the style, or stylodia, and is composed of , the cells of which are receptive to pollen. These may be restricted to the apex of the style or, especially in wind pollinated species, cover a wide surface. The stigma receives pollen and it is on the stigma that the pollen grain germinates. Often sticky, the stigma is adapted in various ways to catch and trap pollen with various hairs, flaps, or sculpturings. The pollen may be captured from the air (wind-borne pollen, anemophily), from visiting insects or other animals ( biotic pollination), or in rare cases from surrounding water (hydrophily). Stigma can vary from long and sle ...
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Stamen
The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament and an anther which contains ''sporangium, microsporangia''. Most commonly anthers are two-lobed and are attached to the filament either at the base or in the middle area of the anther. The sterile tissue between the lobes is called the connective, an extension of the filament containing conducting strands. It can be seen as an extension on the dorsal side of the anther. A pollen grain develops from a microspore in the microsporangium and contains the male gametophyte. The stamens in a flower are collectively called the androecium. The androecium can consist of as few as one-half stamen (i.e. a single locule) as in ''Canna (plant), Canna'' species or as many as 3,482 stamens which have been counted in the saguaro (''Carnegiea gigantea'' ...
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