Cladonia Borealis
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Cladonia Borealis
''Cladonia borealis'', commonly known as the boreal cup lichen, is a species of lichen in the genus ''Cladonia''. ---- Description Cladonia borealis is yellowish green to brown in color. It occurs in North and South America, Antarctica, Eurasia and many on islands. The ascoma, when present is apothecial. Biochemistry Its secondary metabolites include 4-O-demethylbarbatic acid, barbatic acid, rhodocladonic acid and usnic acid. See also * List of Cladonia species, List of ''Cladonia'' species References External links North American Lichen ChecklistUSDA Plants Database



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Lichen
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Cladonia
''Cladonia'' is a genus of moss-like lichens in the family Cladoniaceae. They are the primary food source for reindeer/caribou. ''Cladonia'' species are of economic importance to reindeer-herders, such as the Sami in Scandinavia or the Nenets in Russia. Antibiotic compounds are extracted from some species to create antibiotic cream. The light green species ''Cladonia stellaris'' is used in flower decorations. Although the phylogeny of the genus ''Cladonia'' is still under investigation, two main morphological groups are commonly differentiated by taxonomists: the ''Cladonia'' morpho-type and the ''Cladina'' morpho-type. The ''Cladonia'' morpho-type has many more species, and is generally described as a group of squamulose (grow from squamules), cup-bearing lichens. The ''Cladina'' morpho-types are often referred to as forage lichens, mat-forming lichens, or reindeer lichens (due to their importance as caribou winter forage). ''Cladonia perforata'' ("perforate cladonia") is o ...
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Barbatic Acid
Barbatic acid is an organic compound that is made by some lichens. It is in the structural class known as depsides. It is particularly common in the genera ''Usnea'' (the beard lichens) and ''Cladonia''. History The compound was first isolated in 1880 from the lichen '' Usnea barbata'' by chemists John Stenhouse and Charles Groves. The compound coccellic acid, isolated from '' Cladonia coccifera'', was later shown to be the same compound as barbatic acid. Properties Biosynthetically, barbatic acid is made of two units of orsellinate derivatives that are created by an aromatic synthase enzyme. The repeated action of this enzyme produces an 8-carbon polyketide intermediate that is cyclized. Barbatic acid's IUPAC name is 2-hydroxy-4-(2-hydroxy-4-methoxy-3,6-dimethylbenzoyl)oxy-3,6-dimethylbenzoic acid. Its chemical formula is C19H20O7; it has a molecular mass of 360.36  grams per mole. In its purified crystalline form, it exists as various forms: small rhombic prisms, lo ...
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Usnic Acid
Usnic acid is a naturally occurring dibenzofuran derivative found in several lichen species with the formula C18H16O7. It was first isolated by German scientist W. Knop in 1844 and first synthesized between 1933-1937 by Curd and Robertson. Usnic acid was identified in many genera of lichens including ''Usnea'', '' Cladonia'', ''Hypotrachyna'', ''Lecanora'', ''Ramalina'', '' Evernia'', ''Parmelia'' and '' Alectoria''. Although it is generally believed that usnic acid is exclusively restricted to lichens, in a few unconfirmed isolated cases the compound was found in kombucha tea and non-lichenized ascomycetes. At normal conditions, usnic acid is a bitter, yellow, solid substance. It is known to occur in nature in both the d- and l-forms as well as a racemic mixture. Salts of usnic acid are called usnates (e.g. copper usnate). Biological role in lichens Usnic acid is a secondary metabolite in lichens whose role has not been completely elucidated. It is believed that usnic acid prote ...
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List Of Cladonia Species
''Cladonia'' is a large genus of lichens in the family Cladoniaceae. , ''Species Fungorum'' lists 233 species in the genus. A *'' Cladonia acervata'' *'' Cladonia ahtii'' *'' Cladonia alaskana'' *'' Cladonia albofuscescens'' *'' Cladonia albonigra'' *'' Cladonia aleuropoda'' *'' Cladonia alpina'' Yoshim. (1968) *''Cladonia amaurocraea'' *'' Cladonia anaemica'' *'' Cladonia andesita'' *'' Cladonia angustata'' *'' Cladonia appalachensis'' *'' Cladonia apodocarpa'' *''Cladonia arbuscula'' *'' Cladonia archeri'' *'' Cladonia arcuata'' *'' Cladonia argentea'' *'' Cladonia asahinae'' *'' Cladonia atlantica'' A.Evans (1944) *'' Cladonia atrans'' *'' Cladonia attacta'' *'' Cladonia awasthiana'' B *'' Cladonia bacillaris'' *'' Cladonia bacilliformis'' *'' Cladonia bahiana'' *'' Cladonia bangii'' *'' Cladonia bellidiflora'' *'' Cladonia berghsonii'' *'' Cladonia bimberiensis'' *'' Cladonia borbonica'' *'' Cladonia borealis'' *'' Clad ...
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Lichens Described In 1989
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

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Lichen Species
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (