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Cichorioideae
The Cichorioideae are a subfamily of the Family (biology), family Asteraceae of flowering plants. Familiar members of Cichorioideae include lettuce, dandelions, chicory and ''Gazania'' species. The subfamily comprises about 240 genera and about 2900 species. It is heterogeneous and hard to characterize except with molecular characters. Taxonomy The subfamily as understood in 1998 turned out to be paraphyletic, based on Research, studies of DNA sequences, so a number of Tribe (biology), tribes were moved to new subfamilies. Names for the new subfamilies were published in 2002. In 2004, 2007, and 2008, molecular phylogenetic studies further clarified Systematics, relationships within Cichorioideae.Sterling C. Keeley, Zac H. Forsman, and Raymund Chan. 2007. "A phylogeny of the "evil tribe" (Vernonieae: Compositae) reveals Old/New World long distance dispersal: Support from separate and combined congruent datasets (''trn''L-F, ''ndh''F, ITS)". ''Molecular Phylogenetics and Evolution'' ...
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Asteraceae
The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae were first described in the year 1740. The number of species in Asteraceae is rivaled only by the Orchidaceae, and which is the larger family is unclear as the quantity of extant species in each family is unknown. Most species of Asteraceae are annual, biennial, or perennial herbaceous plants, but there are also shrubs, vines, and trees. The family has a widespread distribution, from subpolar to tropical regions in a wide variety of habitats. Most occur in hot desert and cold or hot semi-desert climates, and they are found on every continent but Antarctica. The primary common characteristic is the existence of sometimes hundreds of tiny individual florets which are held together by protective involucres in flower heads, or more t ...
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Platycarpha
''Platycarpha'' is a genus of South African plants dandelion subfamily within the daisy family.Lessing, Christian Friedrich. 1831. Linnaea 6: 688.
in Latin Recent studies have suggested splitting ''Platycarpha'' into two genera, ''Platycarpha'' and '' Platycarphella.'' Further work remains to be done to resolve this.Per Ola Karis. 2007. "Arctotideae" pages 200-207. In: Klaus Kubitzki (series editor); Joachim W. Kadereit and Charles Jeffrey (volume editors). ''The Families and Genera of Vascular Plants'' volume VIII. Springer-Verlag: Berlin; Heidelberg, Germany. David J. Mabberley. 2008. ''Mabberley's Plant-Book'' third edition ...
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Eremothamnus
''Eremothamnus marlothianus'' is a species of shrub in the family Asteraceae, the only species in the genus ''Eremothamnus''. It is native to the coastal desert of Namibia. It is a small shrub with spiny leaves. Taxonomy and systematics The genus ''Eremothamnus'' was erected in 1889 by Otto Hoffmann, when he named its only species, ''Eremothamnus marlothianus''.Otto Hoffmann. 1889. "Plantae Marlothianae. Ein Beitrag zur Kentniss der Flora Südafrikas" ''Botanische Jarhbücher für Systematik, Pflanzengeschichte und Pflanzengeographie'' 10:278. The specific epithet is for Rudolf Marloth (1855–1931), a South African botanist, pharmacist, and analytical chemist. The generic name is derived from the Greek words ''eremos'' and ''thamnos''. Hoffmann did not give an etymology for the name and it has been supposed that it means "solitary shrub",Umberto Quattrocchi. 2000. ''CRC World Dictionary of Plant Names'' volume II, page 929. CRC Press: Boca Raton; New York; Washington,DC; ...
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Hoplophyllum
''Hoplophyllum'' is a genus of flowering plants in the family Asteraceae.Per Ola Karis. 2007. "Arctotideae" pages 200-207. In: Klaus Kubitzki (series editor); Joachim W. Kadereit and Charles Jeffrey (volume editors). ''The Families and Genera of Vascular Plants'' volume VIII. Springer-Verlag: Berlin; Heidelberg, Germany. It has two species, ''Hoplophyllum spinosum'' and ''Hoplophyllum ferox'', both native to South Africa.David J. Mabberley. 2008. ''Mabberley's Plant-Book'' third edition (2008). Cambridge University Press: UK. Both species are shrubs. The leaves are hard and spine-tipped, much longer than wide, and either cylindrical or somewhat flattened. They are grooved with stripes running lengthwise. The type species is ''Hoplophyllum spinosum''.''Hoplophyllum'' In: Index Nominum Genericorum. In: Regnum Vegetabile (see ''External links'' below). ''Hoplophyllum'' is derived from two Greek words, ''hoplon'' "a tool or weapon" and ''phyllon'' "a leaf", a reference to the spi ...
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Warionia
''Warionia'' is a genus in the tribe Cichorieae within the family Asteraceae. The only known species is ''Warionia saharae'', an endemic of Algeria and Morocco, and it is locally known in the Berber language as afessas, abessas or tazart n-îfiss. It is an aromatic, thistle-like shrub of ½–2 m high, that contains a white latex, and has fleshy, pinnately divided, wavy leaves. It is not thorny or prickly. The aggregate flower heads contain yellow disk florets. It flowers from April till June. Because ''Warionia'' is deviant in many respects from any other Asteraceae, different scholars have placed it hesitantly in the Cardueae, Gundelieae, Mutisieae, but now genetic analysis positions it as the sister group to all other Cichorieae. Description ''Wariona'' is an aromatic shrub, usually between ½–2 m, occasionally only 15 cm or up to 3 m high, that has a network of latex-carrying canals throughout the plant with sticky, white, milky latex. It a ...
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Phylogeny
A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological species or other entities based upon similarities and differences in their physical or genetic characteristics. All life on Earth is part of a single phylogenetic tree, indicating common ancestry. In a ''rooted'' phylogenetic tree, each node with descendants represents the inferred most recent common ancestor of those descendants, and the edge lengths in some trees may be interpreted as time estimates. Each node is called a taxonomic unit. Internal nodes are generally called hypothetical taxonomic units, as they cannot be directly observed. Trees are useful in fields of biology such as bioinformatics, systematics, and phylogenetics. ''Unrooted'' trees illustrate only the relatedness of the leaf nodes and do not require the ancestral root ...
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Systematics
Biological systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: cladograms, phylogenetic trees, phylogenies). Phylogenies have two components: branching order (showing group relationships) and branch length (showing amount of evolution). Phylogenetic trees of species and higher taxa are used to study the evolution of traits (e.g., anatomical or molecular characteristics) and the distribution of organisms ( biogeography). Systematics, in other words, is used to understand the evolutionary history of life on Earth. The word systematics is derived from the Latin word ''systema,'' which means systematic arrangement of organisms. Carl Linnaeus used 'Systema Naturae' as the title of his book. Branches and applications In the study of biological systematics, researchers use the different branches to further understand the relations ...
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Incertae Sedis
' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty at specific taxonomic levels is indicated by ' (of uncertain family), ' (of uncertain suborder), ' (of uncertain order) and similar terms. Examples *The fossil plant '' Paradinandra suecica'' could not be assigned to any family, but was placed ''incertae sedis'' within the order Ericales when described in 2001. * The fossil '' Gluteus minimus'', described in 1975, could not be assigned to any known animal phylum. The genus is therefore ''incertae sedis'' within the kingdom Animalia. * While it was unclear to which order the New World vultures (family Cathartidae) should be assigned, they were placed in Aves ''incertae sedis''. It was later agreed to place them in a separate order, Cathartiformes. * Bocage's longbill, ''Motacilla boc ...
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Heterolepis
''Heterolepis'' is a genus of flowering plants in the sunflower family.Per Ola Karis. 2007. "Arctotideae" pages 200-207. In: Klaus Kubitzki (series editor); Joachim W. Kadereit and Charles Jeffrey (volume editors). ''The Families and Genera of Vascular Plants'' volume VIII. Springer-Verlag: Berlin; Heidelberg, Germany. It has threeDavid J. Mabberley. 2008. ''Mabberley's Plant-Book'' third edition (2008). Cambridge University Press: UK. or four species, all endemic to the Western Cape Province in South Africa. ''Heterolepis'' species are shrublets, typically sprawling and about 30 cm high with moderately large yellow flowers. The flowerheads are solitary with glandular peduncles, the surrounding green bracts having membranous margins, especially in the inner rows. The ray florets are female, with a thread-like lobe opposite the ray. The disk florets are bisexual. The seeds (properly speaking the fruits) are flask-shaped, silky, with a pappus of two unequal rows of bris ...
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Cichorieae
The Cichorieae (also called Lactuceae) are a tribe in the plant family Asteraceae that includes 93 genera, more than 1,600 sexually reproductive species and more than 7,000 apomictic species. They are found primarily in temperate regions of the Eastern Hemisphere. Cichorieae all have milky latex and flowerheads that only contain one type of floret. The genera ''Gundelia'' and ''Warionia'' only have disk florets, while all other genera only have ligulate florets. The genera that contain most species are ''Taraxacum'' (Crepidinae subtribe) with about 1,600 apomictic species, ''Hieracium'' with about 770 sexually reproducing and 5,200 apomictic species, and ''Pilosella'' with 110 sexually reproducing and 700 apomictic species (both Hieraciinae). Well-known members include lettuce, chicory, dandelion, and salsify. Description Most species are herbaceous, perennial, short-lived or annual plants, rarely subshrubs, shrubs or vines. All Cichorieae-species have latex canals in both t ...
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