Charaxes Antiquus
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Charaxes Antiquus
''Charaxes antiquus'' is a butterfly in the family Nymphalidae. It is found on the island of São Tomé. The habitat consists of forests and woodland. The species was named by James John Joicey and George Talbot in 1926. Description The butterfly has black edges with two pointy edges on the bottom end, it has a white striped nearly in the middle, and the remainder is colored dark brown and is lighter on the bottom. It is measured 10 centimeters long. Subspecies The species is sometimes treated as a subspecies of ''Charaxes brutus''. Related species Historical attempts to assemble a cluster of presumably related species into a "''Charaxes jasius'' Group" have not been wholly convincing. More recent taxonomic revision, corroborated by phylogenetic research, allow a more rational grouping congruent with cladistic relationships. Within a well-populated clade of 27 related species sharing a common ancestor approximately 16 mya during the Miocene, 26 are now considered together as T ...
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James John Joicey
James John Joicey FES (28 December 1870 – 10 March 1932) was an English amateur entomologist, who assembled an extensive collection of Lepidoptera in his private research museum, called the Hill Museum, in Witley, Surrey. His collection, 40 years in the making, was considered to have been the second largest in the world held privately and to have numbered over 1.5 million specimens. Joicey was a fellow of the Zoological Society of London, the Royal Geographical Society, the Royal Entomological Society, the Royal Horticultural Society, and the Linnean Society of London. Joicey employed specialist entomologists including George Talbot to curate his collection and financed numerous expeditions throughout the world to obtain previously unknown varieties. More than 190 scientific articles were produced during the active period of the Hill Museum. This body of research was described as "a contribution to the study of the exotic Lepidoptera of very great scient ...
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Monophyletic
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic groups are typically characterised by shared derived characteristics ( synapomorphies), which distinguish organisms in the clade from other organisms. An equivalent term is holophyly. The word "mono-phyly" means "one-tribe" in Greek. Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic group'' consists of all of the descendants of a common ancestor minus one or more monophyletic groups. A '' polyphyletic group'' is characterized by convergent features or habits of scientific interest (for example, night-active primates, fruit trees, aquatic insects). The features by which a polyphyletic group is differentiated from others are not inherited from a common ancestor. These definitions have tak ...
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Charaxes
The rajah and pasha butterflies, also known as emperors in Africa and Australia, (genus ''Charaxes'') make up the huge type genus of the brush-footed butterfly subfamily Charaxinae, or leafwing butterflies. They belong to the tribe Charaxini, which also includes the nawab butterflies ('' Polyura''). ''Charaxes'' are tropical Old World butterflies, with by far the highest diversity in sub-Saharan Africa, a smaller number from South Asia to Melanesia and Australia, and a single species ('' C. jasius'') in Europe. They are generally strong flyers and very popular among butterfly collectors. Etymology ''Charaxes'' means "to sharpen" or "to make pointed", referring to the pointed 'tails' on the hind wing. ''Charaxes'' may also be related to ''charax'', meaning 'a sharp stake', or ''charaxis'', a 'notch' or 'incision', which are also features of the hind wing. Biology ''Charaxes'' frequent sunny forest openings and glades where they rest with open or partly open wings sunning themsel ...
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Butterflies Described In 1926
Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group comprises the large superfamily Papilionoidea, which contains at least one former group, the skippers (formerly the superfamily "Hesperioidea"), and the most recent analyses suggest it also contains the moth-butterflies (formerly the superfamily "Hedyloidea"). Butterfly fossils date to the Paleocene, about 56 million years ago. Butterflies have a four-stage life cycle, as like most insects they undergo complete metamorphosis. Winged adults lay eggs on the food plant on which their larvae, known as caterpillars, will feed. The caterpillars grow, sometimes very rapidly, and when fully developed, pupate in a chrysalis. When metamorphosis is complete, the pupal skin splits, the adult insect climbs out, and after its wings have expanded and dried, it flie ...
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Consortium For The Barcode Of Life
The Consortium for the Barcode of Life (CBOL) was an international initiative dedicated to supporting the development of DNA barcoding as a global standard for species identification. CBOL's Secretariat Office is hosted by the National Museum of Natural History, Smithsonian Institution, in Washington, DC. Barcoding was proposed in 2003 by Prof. Paul Hebert of the University of Guelph in Ontario as a way of distinguishing and identifying species with a short standardized gene sequence. Hebert proposed the 658 bases of the Folmer region of the mitochondrial gene cytochrome-C oxidase-1 as the standard barcode region. Hebert is the Director of the Biodiversity Institute of Ontario, the Canadian Centre for DNA Barcoding, and the International Barcode of Life Project (iBOL), all headquartered at the University of Guelph. The Barcode of Life Data Systems (BOLD) is also located at the University of Guelph. CBOL was created in May 2004 with support of the Alfred P. Sloan Foundation, f ...
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Charaxes Andara
''Charaxes andara'' is a butterfly in the family Nymphalidae. It is found in eastern and southern Madagascar, where it is found in Afrotropical forests. It is very similar to ''Charaxes brutus'', of which it has been considered a subspecies. Description Male Edges of abdominal segments above sometimes slightly grey. Wings above with a violet tint; band conspicuously edged with pale blue from M2 of forewing to (SM1) of the hindwing. Forewing; band 6 to 8 mm broad before SM2 (inclusive of blue border), patch M1-M2 3 to 4 mm wide, spot R2-R3 generally elongate, narrow, sometimes prolonged to bar D, but its proximal portion then shaded with brown, spot R1-R2 often also elongate, spots SC4-R1 mostly bluish, small, often only vestigial; marginal spots white as a rule, often slightly orange proximally, in one specimen (in Henley Grose-Smith's collection) all orange and rather larger than usually. Hindwing: two submarginal dots M2-SM2, blue; admarginal bluish-creamy white lin ...
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Charaxes Junius
''Charaxes junius'' is a butterfly in the family Nymphalidae. It is found in Ethiopia and Sudan. The habitat consists of forests and woodland savanna. Description ''Charaxes junius'' is distinguished from ''brutus'' by having the discal band on the upper surface light yellow and in cellule 1 b of the forewing 6-1 mm. in breadth. Abyssinia. - — ''somalicus'' Rothsch. only differs from ''junius'' in the somewhat narrower median band on both wings. Somaliland. Subspecies *''Charaxes junius junius'' (western and south-western Ethiopia) *''Charaxes junius somalicus'' Rothschild, 1900 (southern Ethiopia, south-eastern Sudan) Taxonomy ''Charaxes junius'' Oberthür, 1880 is treated as a subspecies of ''brutus'' (Cramer, 1779) by Van Someren Henning treats ''junius'' as a distinct species on the authority of Plantrou (1983). Plantrou , J. 1983. Systematique biogeographique et evolution des Charaxes Africains (Lepidoptera, Nymphalidae). ''Publications du Laboratoire de Zoologie, Ecole ...
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Pliocene
The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58See the 2014 version of the ICS geologic time scale
million years ago. It is the second and most recent epoch of the Neogene Period in the . The Pliocene follows the Epoch and is followed by the Epoch. Prior to the 2009 ...
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Charaxes Jasius
''Charaxes jasius'', the two-tailed pasha, is a butterfly in the family Nymphalidae. It is the only European species of the genus ''Charaxes''. Divergence of the Mediterranean species ''C. jasius'' from the last common ancestor it shared with its closest related species still flying in the Afrotropical realm most probably occurred around 2 mya, i.e. during the Pliocene. *''Charaxes legeri'' *''Charaxes saturnus'' tat.rev.2005ref name="Turlin 2005"/> *'' Charaxes pelias'' *''Charaxes castor'' *''Charaxes hansali'' Clade 2: contains the three well-populated additional subgroups (19 species) of the jasius Group: *the ''brutus'' subgroup (4 species): *''Charaxes brutus'' *''Charaxes antiquus'' *''Charaxes junius'' *''Charaxes andara'' *the ''pollux'' subgroup (4 species): *'' Charaxes pollux'' *'' Charaxes phoebus'' *'' Charaxes ansorgei'' *'' Charaxes dowsetti'' *the ''eudoxus'' subgroup (11 species): *'' Charaxes eudoxus'' *'' Charaxes lucyae'' *'' Charaxes richelmanni'' *'' Char ...
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George Talbot (entomologist)
George Talbot FES (26 October 1882 – 13 April 1952) was an English entomologist who specialised in butterflies. He wrote about 150 scientific papers, the majority being primarily systematic, consisting of the description of new species or the revision of various genera. He was also responsible for the curation and preservation of the Joicey collection of Lepidoptera prior to its accession by the Natural History Museum. Life and career George Talbot was born "in rather humble circumstances" in Croydon, Surrey, in 1882. As a young man, he was assistant to Percy Ireland Lathy. He then curated for the wealthy amateur butterfly collector Herbert Adams, followed by the insect dealer William Frederick Henry Rosenberg. During the First World War he worked with Arthur Bacot at the Lister Institute on trench fever and typhus diseases carried by lice. From 1915, he was head curator of the large and increasing collection of amateur lepidopterist James John Joicey at the Hill Museu ...
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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