Charaxes Andara
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''Charaxes andara'' is a
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
in the family
Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilionoidea, they are usually medium-sized to large butterflies. Most species have a red ...
. It is found in eastern and southern
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
,Afrotropical Butterflies: File H - Charaxinae - Tribe Charaxini
/ref> where it is found in Afrotropical forests. It is very similar to ''
Charaxes brutus __NOTOC__ ''Charaxes brutus'', the white-barred emperor or white-barred Charaxes, is a butterfly of the family Nymphalidae. It is found in Africa (see subspecies section for detailed information). Its flight period is year-round. Notes on the ...
'', of which it has been considered a subspecies.


Description


Male

Edges of abdominal segments above sometimes slightly grey. Wings above with a violet tint; band conspicuously edged with pale blue from M2 of forewing to (SM1) of the hindwing. Forewing; band 6 to 8 mm broad before SM2 (inclusive of blue border), patch M1-M2 3 to 4 mm wide, spot R2-R3 generally elongate, narrow, sometimes prolonged to bar D, but its proximal portion then shaded with brown, spot R1-R2 often also elongate, spots SC4-R1 mostly bluish, small, often only vestigial; marginal spots white as a rule, often slightly orange proximally, in one specimen (in
Henley Grose-Smith Henley Grose-Smith (1833–1911) was an English entomologist who specialised in Lepidoptera. Grose-Smith described many new taxa of butterflies from his own collections and those of Walter Rothschild. His collections were sold to James John Joicey ...
's collection) all orange and rather larger than usually. Hindwing: two submarginal dots M2-SM2, blue; admarginal bluish-creamy white line seldom continuous, mostly very widely interrupted between veins, there remaining only triangular spots at the ends of the veins, these spots extending into tails, anal admarginal spot, however, linear and transverse, bluish olive; discal band stopping at (SM1). Underside rufous red from base to discal band, bars without grey centres. Forewing: cell-bar 3 somewhat rounded, 4 rather slender, as is bar D; median bars SC3-R2 also slender, bars SC3-R2 each very much longer than broad (a part of white border) discal bars SC4-R1 dark olive; outer area of wing from white band to margin tawny ochraceous; black postdiscal patches R1-M! rather smaller than in the continental forms, less pointed. Hindwing: basal costal bar not touching white costal edge, often rounded, almost circular, white band posteriorly as well as anteriorly with brown spots or vestiges of such; interspace between white discal band and black postdiscal bars cinnamon rufous, as a rule darker than basal area; postdiscal bars C-M2 transverse, not or slightly curved, rather thin, not dilated in middle, edged olive distally; submarginal interspace chestnut, seldom rufous chestnut, much wider than in the other forms, bar R1-R2 5 mm distant from submarginal bar, much shaded with white distally, interspaces M1-SM2 olive, with blue and white submarginal dots; submarginal bars blackish olive, transverse, linear; wing more heavily dentate than in the continental forms of ''C. brutus'', tail R3 7 to 8 mm long, tail M1 3 to 5 mm, tail M2 to  mm.


Female

Wings above less bluish than in male. Forewing: costal margin more or less tawny; band pale orange, white from M2 backwards, posteriorly edged with pale blue at both sides, but especially proximally, 10 mm broad at SM2, patch R3-M1 7 mm long, oblique proximally, patch R2-R3 extended to bar D, patch R1-R2 about twice as long as broad, with two discal spots in front, besides the two rounded postdiscal spots SC1-R1, discal spot SC5-R1 rather larger, produced distad along R1 and mostly joined to the postdiscal spot, discal spot SC4=SC5 linear, an orange streak also in front of SC4; marginal spots pale orange. Hindwing: band broadly bordered with bluish white proximally, basal half of abdominal fold also milky white, pale blue distal border of band gradually narrowing costad; blue submarginal dots M2-SM2 heavy, fused together as a rule, dot M1-M2 often present; admarginal line orange between, creamy buff or bluish at veins, nearly continuous, or interrupted at the veins, often obsolete from C to R1, anal portion as in male but wider. Underside: as in male, but submarginal area of hindwing more olive and white, the chestnut scaling very much reduced, sometimes almost entirely replaced by olive and white; tails and teeth as in male, rather broader at base and longer. Length of forewing: male 34–41 mm, female 42–46 mm. Habitat: Madagascar, apparently all over the island in suitable localities; a forest insect.


Related species

Historical attempts to assemble a cluster of presumably related species into a "''Charaxes jasius'' Group" have not been wholly convincing. More recent taxonomic revision, corroborated by phylogenetic research, allow a more rational grouping congruent with cladistic relationships. Within a well-populated
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
of 27 related species sharing a common ancestor approximately 16 mya during the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
, 26 are now considered together as The ''jasius'' Group. One of the two lineages within this clade forms a robust
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
group of seven species sharing a common ancestor approximately 2-3 mya, i.e. during the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58"Out of Africa again: A phylogenetic hypothesis of the genus ''Charaxes (Lepidoptera: Nymphalidae)'' based on five gene regions"
Aduse-Poku, Vingerhoedt, Wahlberg. Molecular Phylogenetics and Evolution (2009) 53;463–478
and are considered as the ''jasius'' subgroup. The second lineage leads to 19 other species within the Jasius group, which are split in to three well-populated subgroups of closely related species. The ''jasius'' Group (26 Species). Clade 1: the ''jasius'' subgroup. Clade 2: contains the three well-populated additional subgroups (19 species) of the jasius Group, called the ''brutus'', ''pollux'', and ''eudoxus'' subgroups. *the ''brutus'' subgroup (4 Species) *''
Charaxes brutus __NOTOC__ ''Charaxes brutus'', the white-barred emperor or white-barred Charaxes, is a butterfly of the family Nymphalidae. It is found in Africa (see subspecies section for detailed information). Its flight period is year-round. Notes on the ...
'' *''
Charaxes antiquus ''Charaxes antiquus'' is a butterfly in the family Nymphalidae. It is found on the island of São Tomé. The habitat consists of forests and woodland. The species was named by James John Joicey and George Talbot in 1926. Description The butter ...
'' *''
Charaxes junius ''Charaxes junius'' is a butterfly in the family Nymphalidae. It is found in Ethiopia and Sudan. The habitat consists of forests and woodland savanna. Description ''Charaxes junius'' is distinguished from ''brutus'' by having the discal band on ...
'' *''Charaxes andara'' Further exploration of the phylogenetic relationships amongst existing ''Charaxes'' taxa is required to improve clarity.


References

*
Victor Gurney Logan Van Someren Victor Gurney Logan Van Someren (1886 in Melbourne – 24 July 1976) was a zoologist and entomologist. Van Someren was born in Australia. He attended George Watson's College and studied zoology at University of Edinburgh. He was also a dentist ...
, 1970 Revisional notes on African ''Charaxes'' (Lepidoptera: Nymphalidae). Part VI. ''Bulletin of the British Museum'' (Natural History) (Entomology) 197-25


External links


Butterfly corner
Images from Naturhistorisches Museum, Naturhistorisches Museum Wien
''Charaxes andara'' images
at
Consortium for the Barcode of Life The Consortium for the Barcode of Life (CBOL) was an international initiative dedicated to supporting the development of DNA barcoding as a global standard for species identification. CBOL's Secretariat Office is hosted by the National Museum of ...

African Butterfly Database
Range map via search {{Taxonbar, from=Q5073964 Butterflies described in 1873
andara Andara is a village in Mukwe Constituency in the Kavango East region of north-eastern Namibia. Located east of Rundu, it is inhabited primarily by the Hambukushu people. Founding of the Catholic mission Catholic fathers of the organization Mi ...
Endemic fauna of Madagascar Butterflies of Africa Taxa named by Christopher Ward (entomologist)