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Chalicotheres
Chalicotheres (from Greek language, Greek ''wikt:χάλιξ, chalix'', "gravel" and ''wikt:θηρίον, therion'', "beast") are an extinct clade of herbivorous, odd-toed ungulate (perissodactyl) mammals that lived in North America, Eurasia, and Africa from the Eocene, Middle Eocene until the Early Pleistocene, existing from 48.6 to 1.806 Annum, mya. They are one of the five major Evolutionary radiation, radiations of perissodactyls, with three groups living (Equidae, horses, plus the extinct Palaeotheriidae, paleotheres; Rhinoceros, rhinoceroses; Tapiroidea, tapirs), and two extinct (Brontotheriidae, brontotheres and chalicotheres). Description Unlike modern perissodactyls, chalicotheres had clawed feet. They had longer forelimbs and shorter hind limbs, lower incisors that cropped food against a toothless pad in the upper jaw, low-crowned molar teeth, and were browsers on trees and shrubs throughout their history. They evolved in two different directions, which became separate ...
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Odd-toed Ungulate
Odd-toed ungulates, mammals which constitute the taxonomic order Perissodactyla (, ), are animals—ungulates—who have reduced the weight-bearing toes to three (rhinoceroses and tapirs, with tapirs still using four toes on the front legs) or one (equines, third toe) of the five original toes. The non-weight-bearing toes are either present, absent, vestigial, or positioned posteriorly. By contrast, the even-toed ungulates bear most of their weight equally on four or two (an even number) of the five toes: their third and fourth toes. Another difference between the two is that odd-toed ungulates digest plant cellulose in their intestines rather than in one or more stomach chambers as even-toed ungulates, with the exception of Suina, do. The order includes about 17 species divided into three families: Equidae (horses, asses, and zebras), Rhinocerotidae (rhinoceroses), and Tapiridae (tapirs). Despite their very different appearances, they were recognized as related families in ...
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Chalicotherium
''Chalicotherium'' (Ancient Greek /, -: pebble/gravel + /, diminutive of / : beast) is a genus of extinct odd-toed ungulates of the order Perissodactyla and family Chalicotheriidae. The genus is known from Europe and Asia, from the Middle Miocene to Late Miocene. This animal would look much like other chalicotheriid species: an odd-looking herbivore with long clawed forelimbs and stouter weight-bearing hindlimbs. The type species, ''Chalicotherium goldfussi'', from Late Miocene Europe, was described by Johann Jakob Kaup in 1833. When the French naturalist George Cuvier first received a cleft claw from Eppelheim, Germany, he identified it as the toe bone of a gigantic pangolin. Description ''Chalicotherium'', like many members of Perissodactyla, was adapted to browsing, though the chalicotheres were uniquely adapted to do so among ungulates. Its arms were long and heavily clawed, allowing them to walk on their knuckles only. The arms were used to reach for the branches ...
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Schizotheriinae
Schizotheriines are one of the two subfamilies of the extinct family Chalicotheriidae, a group of herbivorous odd-toed ungulate (perissodactyl) mammals that lived from the Eocene to the Pleistoscene. The other clade is the Chalicotheriinae. Both clades had claws rather than hooves on their front feet, an adaptation understood as related to feeding. Schizotheriines also had claws on their hind feet. The fossils of both groups are found in environments that had trees and shrubs. While chalicotheriines developed very derived body forms, schizotheriines remained basically similar in shape to other perissodactyls such as horses and tapirs. Like most forest-dwelling ungulates, they had long necks and forelimbs longer than their hindlimbs. Schizotheriines had longer, higher-crowned cheek teeth than chalicotheriines, which indicates they typically fed on tougher vegetation. The sediments where their fossils are found show they also lived in a wider range of environments, from moist forest ...
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Chalicotheriinae
Chalicotheriines are one of the two subfamilies of the extinct family Chalicotheriidae, a group of herbivorous, odd-toed ungulate ''(perissodactyl)'' mammals that lived from the Eocene to the Pleistocene. The other subfamily is the Schizotheriinae. Chalcotheriines evolved unique characteristics for ungulates, with very long forelimbs, short hindlimbs, and a relatively gorilla-like physique, including knuckle-walking on their flexible forelimbs, which bore long curved claws. Members of this subfamily possessed some of the longest forelimbs and shortest hindlimbs in relation to each other out of all extinct animals. Analysis of dental wear implies that most chalicotheriines fed on seeds and fruit. Their claws were likely used in a hook-like manner to pull down branches, suggesting they lived as bipedal browsers. Presence of chalicothere fossils is generally regarded as an indicator of forested environments. Unlike schizotheriines, chalicotheriines were typically confined to moist ...
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Moropus
''Moropus'' (meaning "slow foot") is an extinct genus of large perissodactyl ("odd-toed" ungulate) mammal in the chalicothere family. They were endemic to North America during the Miocene from ~20.4—13.6  Mya, existing for approximately . ''Moropus'' belonged to the schizotheriine subfamily of chalicotheres, and has the best fossil record of any member of this group; numbers of individuals, including complete skeletons, have been found. The closest extant relatives of ''Moropus'' are other perissodactyls: Equus (genus), horses, Rhinoceros, rhinos, and tapirs. Description Like other chalicothere Chalicotheres (from Greek '' chalix'', "gravel" and '' therion'', "beast") are an extinct clade of herbivorous, odd-toed ungulate (perissodactyl) mammals that lived in North America, Eurasia, and Africa from the Middle Eocene until the Early Plei ...s, ''Moropus'' differed from typical ungulates in having large claws, rather than hooves, on the feet. Three large, highly compress ...
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Brontotheriidae
Brontotheriidae is a family of extinct mammals belonging to the order Perissodactyla, the order that includes horses, rhinoceroses, and tapirs. Superficially, they looked rather like rhinos, although they were actually more closely related to horses; Equidae and Brontotheriidae make up the suborder Hippomorpha. They lived around 56–34 million years ago, until the very close of the Eocene. Characteristics and evolution Brontotheres retain four toes on their front feet and three toes on their hind feet. Their teeth are adapted to shearing (cutting) relatively nonabrasive vegetation. Their molars have a characteristic W-shaped ectoloph (outer shearing blade). The evolutionary history of this group is well known due to an excellent fossil record in North America. The earliest brontotheres, such as ''Eotitanops'', were rather small, no more than a meter in height, and hornless. Brontotheres evolved massive bodies, although some small species such as ''Nanotitanops'' did pers ...
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Giraffe
The giraffe is a large African hoofed mammal belonging to the genus ''Giraffa''. It is the tallest living terrestrial animal and the largest ruminant on Earth. Traditionally, giraffes were thought to be one species, ''Giraffa camelopardalis'', with nine subspecies. Most recently, researchers proposed dividing them into up to eight extant species due to new research into their mitochondrial and nuclear DNA, as well as morphological measurements. Seven other extinct species of ''Giraffa'' are known from the fossil record. The giraffe's chief distinguishing characteristics are its extremely long neck and legs, its horn-like ossicones, and its spotted coat patterns. It is classified under the family Giraffidae, along with its closest extant relative, the okapi. Its scattered range extends from Chad in the north to South Africa in the south, and from Niger in the west to Somalia in the east. Giraffes usually inhabit savannahs and woodlands. Their food source is leaves, frui ...
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Tapiroidea
Tapiroidea is a superfamily of perissodactyls which includes the modern tapir. Members of the superfamily are small to large browsing mammals, roughly pig-like in shape, with short, prehensile snouts. Their closest relatives are the other odd-toed ungulates, including horses and rhinoceroses. Taxonomically, they are placed in suborder Ceratomorpha along with the rhino superfamily, Rhinocerotoidea. The first members of Tapiroidea appeared during the Early Eocene, 55 million years ago. Evolution and natural history The first tapirids, such as ''Heptodon'', appeared in the early Eocene. They appeared very similar to modern forms, but were about half the size, and lack the proboscis. The first true tapirs, appeared in the Oligocene, and by the Miocene, such genera as ''Miotapirus'' were almost indistinguishable to the extant species. It is believed that Asian and American tapirs diverged around 20 to 30 million years ago, and that tapirs migrated from North America and Central Ameri ...
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Eurasia
Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago and the Russian Far East to the east. The continental landmass is bordered by the Atlantic Ocean and Africa to the west, the Pacific Ocean to the east, the Arctic Ocean to the north, and by Africa, the Mediterranean Sea, and the Indian Ocean to the south. The division between Europe and Asia as two continents is a historical social construct, as many of their borders are over land; thus, in some parts of the world, Eurasia is recognized as the largest of the six, five, or four continents on Earth. In geology, Eurasia is often considered as a single rigid megablock. However, the rigidity of Eurasia is debated based on paleomagnetic data. Eurasia covers around , or around 36.2% of the Earth's total land area. It is also home to the largest ...
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Rhinoceros
A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea.) Two of the extant species are native to Africa, and three to South and Southeast Asia. Rhinoceroses are some of the largest remaining megafauna: all weigh at least one tonne in adulthood. They have a herbivorous diet, small brains (400–600 g) for mammals of their size, one or two horns, and a thick (1.5–5 cm), protective skin formed from layers of collagen positioned in a lattice structure. They generally eat leafy material, although their ability to ferment food in their hindgut allows them to subsist on more fibrous plant matter when necessary. Unlike other perissodactyls, the two African species of rhinoceros lack teeth at the front of their mouths; they rely instead on their lips to pl ...
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Palaeotheriidae
Palaeotheriidae is an extinct family of herbivorous perissodactyl mammals related to equids. They ranged across Europe and Asia from the Eocene through to the early Oligocene 55–33  Ma, existing for approximately . Living in dense forests, they ate soft leaves, shoots, berries, and leaf matter picked up from the forest floor. Morphology Palaeothere sizes ranged from at the shoulder, and weighed an estimated .S. Legendre. 1988. Les communautes de mammiferes du Paleogene (Eocene superieur et Oligocene) d'Europe occidentale: structure, milieux et evolution. Ph.D. thesis, Universite des Sciences et Techniques du Languedoc, Montpellier, France. 2 volumes. 1-265 Extinction Evidence suggests that the palaeotheriidae went extinct in Eurasia during the Early Oligocene, approximately 33  Ma, as part of a faunal turnover event known as the Grande Coupure. The Eocene-Oligocene transition marked a significant global cooling event caused by the onset of Antarctic glaciation. ...
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