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Chaetothyriomycetidae
Chaetothyriomycetidae is a subclass of ascomycete within the class Eurotiomycetes. Many species in Chaetothyriomycetidae are lichens. Morphology Chaetothyriomycetidae produce a cleistothecium An ascocarp, or ascoma (), is the fruiting body ( sporocarp) of an ascomycete phylum fungus. It consists of very tightly interwoven hyphae and millions of embedded asci, each of which typically contains four to eight ascospores. Ascocarps are ... through which they distribute their spores. Gallery File:Exophiala phaeomuriformis.jpg File:Chromoblastomycosis 2.jpg File:Fonsecaea pedrosoi PHIL 2920 lores.jpg File:Fonsecaea pedrosoi.jpg File:Cuerpo Esclerotico.JPG File:Fonsecaea.JPG File:Phialophora verrucosa 002.jpg File:A lichen - Verrucaria mucosa - geograph.org.uk - 1055602.jpg File:Wahlenbergiella mucosa Jymm.jpg File:Verrucaria ochrostoma (Verrucariaceae), Renkum, the Netherlands.jpg File:Verrucaria muralis (Verrucariaceae), Renkum, the Netherlands.jpg File:Verrucaria macrosto ...
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Eurotiomycetes
Eurotiomycetes is a large class of ascomycetes with cleistothecial ascocarps within the subphylum Pezizomycotina, currently containing around 3810 species according to the Catalogue of Life. It is the third largest lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship. It contains most of the fungi previously known morphologically as "Plectomycetes".


Systematics and phylogeny


Internal relationships

The class Eurotiomycetes was circumscription (taxonomy), circumscribed in 1997 by Sweden, Swedish mycologists Ove Erik Eriksson and Katarina Winka. At that time ...

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Chaetothyriales
The Chaetothyriales are an order of ascomycetous fungi in the class Eurotiomycetes and within the subclass Chaetothyriomycetidae. The order was circumscribed in 1987 by mycologist Margaret Elizabeth Barr-Bigelow. Families and genera , Species Fungorum includes 9 families, 97 genera, and 691 species in the Chaetothyriales. The following list shows the families, genera, and number of species in the Chaetothyriales, adapted from a recent (2020) taxonomic and nomenclatural review of the order. * Chaetothyriaceae ::'' Actinocymbe'' – 3 spp. ::'' Aithaloderma'' – 12 spp. ::'' Aphanophora'' – 1 sp. ::'' Arthrophiala'' – 1 sp. ::'' Camptophora'' – 2 spp. ::'' Ceramothyrium'' – 39 spp. ::'' Ceratocarpia'' – 3 spp. ::'' Chaetothyriomyces'' – 1 sp. ::'' Chaetothyrium'' – 67 spp. ::'' Cyphellophoriella'' – 1 sp. ::'' Euceramia'' – 3 spp. ::'' Longihyalospora'' – 2 spp. ::'' Microcallis'' – 9spp. ::'' Nullicamyces'' – 1 sp. ::'' Phaeosaccardinula'' – 41 spp. :: ...
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Verrucariales
Verrucariales is an order of ascomycetous fungi within the subclass Chaetothyriomycetidae of the class Eurotiomycetes. Although most of the Verrucariales are lichenised, the family Sarcopyreniaceae consists of 11 species of lichenicolous (lichen-dwelling) fungi. Phylogenomic analysis suggests that the divergence between the lichenized Verrucariales and nonlichenized Chaetothyriales The Chaetothyriales are an order of ascomycetous fungi in the class Eurotiomycetes and within the subclass Chaetothyriomycetidae. The order was circumscribed in 1987 by mycologist Margaret Elizabeth Barr-Bigelow. Families and genera , Species ... occurred about 131 million years ago. Genera of uncertain placement There are some genera in the Verrucariales that have not been placed with certainty into any family. These are: *'' Botryolepraria'' – 2 spp. *'' Gemmaspora'' – 1 sp. *'' Kalbiana'' – 1 sp. *'' Merismatium'' – 10 spp. References Ascomycota orders Lichen orders ...
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Pyrenocollema Halodytes
''Pyrenocollema'' is a genus of lichenized fungi in the biological division Ascomycota. Its relationship to other genera is unclear, and it has not been assigned to any family, order or class Class or The Class may refer to: Common uses not otherwise categorized * Class (biology), a taxonomic rank * Class (knowledge representation), a collection of individuals or objects * Class (philosophy), an analytical concept used differentl .... Species *'' Pyrenocollema elegans'' *'' Pyrenocollema halodytes'' *'' Pyrenocollema occidentalipamiricum'' *'' Pyrenocollema orustense'' *'' Pyrenocollema pelvetiae'' *'' Pyrenocollema tichothecioides'' *'' Pyrenocollema tremelloides'' References Xanthopyreniaceae Lichen genera Ascomycota genera Taxa described in 1895 {{Ascomycota-stub ...
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Littoraria Irrorata
''Littoraria irrorata'', also known by the common name the marsh periwinkle, is a species of sea snail, a marine gastropod mollusk in the family Littorinidae. The specific epithet ' means 'moistened' or 'dewy.' This species occurs in salt marshes on the Atlantic coast and Gulf Coast of North America, from Massachusetts to Texas. Some colonies of this species of snail are the only mollusks known to practice fungiculture. ''L. irrorata'' is an essential part of the salt marsh ecosystem. This is displayed in its strong relationship with ''Sporobolus alterniflorus'', also known as ''Spartina alterniflora'', a grass commonly found in abundance in salt marshes. Description The maximum recorded shell length is . ''L. irrorata'' is extremely temperature tolerant. The snail has the ability to retract its foot into its shell when experiencing thermal stress which allows them to avoid water loss by evaporation and survive in high temperatures. Distribution This species can be found along ...
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Alexander Borissovitch Doweld
Alexander is a male given name. The most prominent bearer of the name is Alexander the Great, the king of the Ancient Greek kingdom of Macedonia who created one of the largest empires in ancient history. Variants listed here are Aleksandar, Aleksander and Aleksandr. Related names and diminutives include Iskandar, Alec, Alek, Alex, Alexandre, Aleks, Aleksa and Sander; feminine forms include Alexandra, Alexandria, and Sasha. Etymology The name ''Alexander'' originates from the (; 'defending men' or 'protector of men'). It is a compound of the verb (; 'to ward off, avert, defend') and the noun (, genitive: , ; meaning 'man'). It is an example of the widespread motif of Greek names expressing "battle-prowess", in this case the ability to withstand or push back an enemy battle line. The earliest attested form of the name, is the Mycenaean Greek feminine anthroponym , , (/Alexandra/), written in the Linear B syllabic script. Alaksandu, alternatively called ''Alakasandu'' or ' ...
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Pyrenulales
The ''Pyrenulales'' are an order of ascomycetous fungi within the class Eurotiomycetes and within the subphylum Pezizomycotina. Taxonomy As of 2022 the order contains one family, 14 genera and around 296 species. *Order Pyrenulales **Family Pyrenulaceae ***'' Anthracothecium'' – 5 species ***'' Blastodesmia'' – 1 species ***'' Clypeopyrenis'' – 2 species ***'' Distopyrenis'' – 8 species ***'' Granulopyrenis'' – 6 species ***'' Lithothelium'' – 28 species ***'' Mazaediothecium'' – 4 species ***'' Pyrenographa'' – 1 species ***'' Pyrenowilmsia'' – 1 species ***'' Pyrenula'' (='' Heufleridium'' ; ='' Stromatothelium'' ) – circa 225 species ***'' Pyrgillus'' – 8 species ***'' Sulcopyrenula'' – 5 species **Pyrenulales incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of ...
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Class (biology)
In biological classification, class ( la, classis) is a taxonomic rank, as well as a taxonomic unit, a taxon, in that rank. It is a group of related taxonomic orders. Other well-known ranks in descending order of size are life, domain, kingdom, phylum, order, family, genus, and species, with class fitting between phylum and order. History The class as a distinct rank of biological classification having its own distinctive name (and not just called a ''top-level genus'' ''(genus summum)'') was first introduced by the French botanist Joseph Pitton de Tournefort in his classification of plants that appeared in his ''Eléments de botanique'', 1694. Insofar as a general definition of a class is available, it has historically been conceived as embracing taxa that combine a distinct ''grade'' of organization—i.e. a 'level of complexity', measured in terms of how differentiated their organ systems are into distinct regions or sub-organs—with a distinct ''type'' of construction, ...
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Ascomycota
Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The defining feature of this fungal group is the " ascus" (), a microscopic sexual structure in which nonmotile spores, called ascospores, are formed. However, some species of the Ascomycota are asexual, meaning that they do not have a sexual cycle and thus do not form asci or ascospores. Familiar examples of sac fungi include morels, truffles, brewers' and bakers' yeast, dead man's fingers, and cup fungi. The fungal symbionts in the majority of lichens (loosely termed "ascolichens") such as ''Cladonia'' belong to the Ascomycota. Ascomycota is a monophyletic group (it contains all descendants of one common ancestor). Previously placed in the Deuteromycota along with asexual species from other fungal taxa, asexual (or anamorphic) ascomyce ...
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Lichen
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (



Ascocarp
An ascocarp, or ascoma (), is the fruiting body ( sporocarp) of an ascomycete phylum fungus. It consists of very tightly interwoven hyphae and millions of embedded asci, each of which typically contains four to eight ascospores. Ascocarps are most commonly bowl-shaped (apothecia) but may take on a spherical or flask-like form that has a pore opening to release spores (perithecia) or no opening (cleistothecia). Classification The ascocarp is classified according to its placement (in ways not fundamental to the basic taxonomy). It is called ''epigeous'' if it grows above ground, as with the morels, while underground ascocarps, such as truffles, are termed ''hypogeous''. The structure enclosing the hymenium is divided into the types described below (apothecium, cleistothecium, etc.) and this character ''is'' important for the taxonomic classification of the fungus. Apothecia can be relatively large and fleshy, whereas the others are microscopic—about the size of flecks ...
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Spore
In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, fungi and protozoa. Bacterial spores are not part of a sexual cycle, but are resistant structures used for survival under unfavourable conditions. Myxozoan spores release amoeboid infectious germs ("amoebulae") into their hosts for parasitic infection, but also reproduce within the hosts through the pairing of two nuclei within the plasmodium, which develops from the amoebula. In plants, spores are usually haploid and unicellular and are produced by meiosis in the sporangium of a diploid sporophyte. Under favourable conditions the spore can develop into a new organism using mitotic division, producing a multicellular gametophyte, which eventually goes on to produce gametes. Two gametes fuse to form a zygote which develops into a new s ...
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