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Centrohelida
The centrohelids or centroheliozoa are a large group of heliozoan protists. They include both mobile and sessile forms, found in freshwater and marine environments, especially at some depth. Characteristics Individuals are unicellular and spherical, usually around 30–80 μm in diameter, and covered with long radial axopods, narrow cellular projections that capture food and allow mobile forms to move about. A few genera have no cell covering, but most have a gelatinous coat holding scales and spines, produced in special deposition vesicles. These may be organic or siliceous and come in various shapes and sizes. For instance, in ''Raphidiophrys'' the coat extends along the bases of the axopods, covering them with curved spicules that give them a pine-treeish look, and in ''Raphidiocystis'' there are both short cup-shaped spicules and long tubular spicules that are only a little shorter than the axopods. Some other common genera include ''Heterophrys'', ''Actinocystis'', and ' ...
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Eukaryota
Eukaryotes () are organisms whose Cell (biology), cells have a cell nucleus, nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the Three-domain system, three domains of life. Bacteria and Archaea (both prokaryotes) make up the other two domains. The eukaryotes are usually now regarded as having emerged in the Archaea or as a sister of the Asgard (archaea), Asgard archaea. This implies that there are only Two-domain system, two domains of life, Bacteria and Archaea, with eukaryotes incorporated among archaea. Eukaryotes represent a small minority of the number of organisms, but, due to their generally much larger size, their collective global biomass (ecology), biomass is estimated to be about equal to that of prokaryotes. Eukaryotes emerged approximately 2.3–1.8 billion years ago, during the Proterozoic eon, likely as Flagellated cell, flagellated phagotrophs. The ...
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Cryptomonads-haptophytes Assemblage
The cryptomonads-haptophytes assemblage is a proposed monophyletic grouping of unicellular eukaryotes that are not included in the SAR supergroup. Several alternative names have been used for the group, including Hacrobia (derived from "ha-" referring to Haptophyta, "-cr-" referring to cryptomonads, and "-bia" as a general suffix referring to life); CCTH (standing for Cryptophyta, Centrohelida, Telonemia and Haptophyta); and "Eukaryomonadae". , it is unclear whether this group is monophyletic or not; results of phylogenetic studies are "often dependent on the selection of taxa and gene data set". Two 2012 studies produced opposite results. Members In the past, heterokonts, haptophytes, and cryptomonads have sometimes been grouped together in a group known as chromists. Though the heterokonts are now split out, Cryptophyta and Haptophyta are considered in some studies to be closely related (and are sometimes simply referred to as the "Cryptophyta+Haptophyta" group). A 2009 pape ...
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Haptista
Haptista is a proposed group of protists made up of centrohelids and haptophytes. Phylogenomic studies indicate that Haptista, together with ''Ancoracysta twista'', forms a sister clade to the SAR+Telonemia supergroup, but it may also be sister to the Cryptista (+Archaeplastida). It is thus one of the earliest diverging Diaphoretickes. Taxonomy Based on studies done by Cavalier-Smith, Chao & Lewis 2015 and Ruggiero et al. 2015. * Subphylum Centroheliozoa Cushman & Jarvis 1929 sensu Durrschmidt & Patterson 1987 eliozoa Haeckel 1862 stat. n. Margulis 1974 em. Cavalier-Smith 2003** Class Centrohelea Kuhn 1926 stat. n. Cavalier-Smith 1993 entroplastiales; Centrohelina Hartmann 1913; Centroplasthelida Febvre-Chevalier 1984* Subphylum Haptophytina Cavalier-Smith 2015 (Haptophyta Hibberd 1976 sensu Ruggerio et al. 2015) ** Clade Rappemonada Kim et al. 2011 *** Class Rappephyceae Rappephyceae, or Rappemonads, are a small family of protists first described in 2011, of uncertain phylog ...
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Hacrobia
The cryptomonads-haptophytes assemblage is a proposed monophyletic grouping of unicellular eukaryotes that are not included in the SAR supergroup. Several alternative names have been used for the group, including Hacrobia (derived from "ha-" referring to Haptophyta, "-cr-" referring to cryptomonads, and "-bia" as a general suffix referring to life); CCTH (standing for Cryptophyta, Centrohelida, Telonemia and Haptophyta); and "Eukaryomonadae". , it is unclear whether this group is monophyletic or not; results of phylogenetic studies are "often dependent on the selection of taxa and gene data set". Two 2012 studies produced opposite results. Members In the past, heterokonts, haptophytes, and cryptomonads have sometimes been grouped together in a group known as chromists. Though the heterokonts are now split out, Cryptophyta and Haptophyta are considered in some studies to be closely related (and are sometimes simply referred to as the "Cryptophyta+Haptophyta" group). A 2009 pape ...
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Raphidiophrys
Raphidiophrys is a genus of centrohelid The centrohelids or centroheliozoa are a large group of heliozoan protists. They include both mobile and sessile forms, found in freshwater and marine environments, especially at some depth. Characteristics Individuals are unicellular and spher ... with radiating axopodia.Zlatogursky, V. V. (2012). ''Raphidiophrys heterophryoidea'' sp. nov. (Centrohelida: Raphidiophryidae), the first heliozoan species with a combination of siliceous and organic skeletal elements. ''European Journal of Protistology, 48''(1), 9-16. doi:10.1016/j.ejop.2011.09.004 ''R. intermedia'' is found in the bottom sludge of freshwater bodies in Canada, Chile, Argentina, Australia, New Zealand, Malaysia, Russia, and central Europe. ''Raphidiophrys'' have bipartite scales are a defining characteristic among species. Differences in type and size of scales are used to differentiate amongst the members of this genus. The genus ''Raphidiophrys'' was discovered in 1867 by W. ...
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Heliozoa
Heliozoa, commonly known as sun-animalcules, are microbial eukaryotes (protists) with stiff arms ( axopodia) radiating from their spherical bodies, which are responsible for their common name. The axopodia are microtubule-supported projections from the amoeboid cell body, and are variously used for capturing food, sensation, movement, and attachment. They are similar to Radiolaria, but they are distinguished from them by lacking central capsules and other complex skeletal elements, although some produce simple scales and spines. They may be found in both freshwater and marine environments. Classification Originally the heliozoa were treated together as a formal taxon Heliozoa or Heliozoea, with the rank of class or phylum, but it has been realised that they are polyphyletic, as the various orders show notable differences and are no longer believed to be descended from a single common ancestor. Instead, "heliozoa" is regarded as a descriptive term applying to various lines of proti ...
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Heterophrys
The centrohelids or centroheliozoa are a large group of heliozoan protists. They include both mobile and sessile forms, found in freshwater and marine environments, especially at some depth. Characteristics Individuals are unicellular and spherical, usually around 30–80 μm in diameter, and covered with long radial axopods, narrow cellular projections that capture food and allow mobile forms to move about. A few genera have no cell covering, but most have a gelatinous coat holding scales and spines, produced in special deposition vesicles. These may be organic or siliceous and come in various shapes and sizes. For instance, in ''Raphidiophrys'' the coat extends along the bases of the axopods, covering them with curved spicules that give them a pine-treeish look, and in ''Raphidiocystis'' there are both short cup-shaped spicules and long tubular spicules that are only a little shorter than the axopods. Some other common genera include ''Heterophrys'', ''Actinocystis'', and ' ...
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Cryptomonad
The cryptomonads (or cryptophytes) are a group of algae, most of which have plastids. They are common in freshwater, and also occur in marine and brackish habitats. Each cell is around 10–50 μm in size and flattened in shape, with an anterior groove or pocket. At the edge of the pocket there are typically two slightly unequal flagella. Some may exhibit mixotrophy. Characteristics Cryptomonads are distinguished by the presence of characteristic extrusomes called ejectosomes, which consist of two connected spiral ribbons held under tension. If the cells are irritated either by mechanical, chemical or light stress, they discharge, propelling the cell in a zig-zag course away from the disturbance. Large ejectosomes, visible under the light microscope, are associated with the pocket; smaller ones occur underneath the periplast, the cryptophyte-specific cell surrounding. Except for the class ''Goniomonadea'', which lacks plastids entirely, and ''Cryptomonas paramecium'' (previo ...
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Raphidiophryidae
Raphidiophryidae is a family of mostly freshwater centrohelids. It is the sister family of Acanthocystidae, sharing the trait of presenting silica scales and comprising the clade Chalarothoracina. Two genera, ''Raphidiophrys Raphidiophrys is a genus of centrohelid with radiating axopodia.Zlatogursky, V. V. (2012). ''Raphidiophrys heterophryoidea'' sp. nov. (Centrohelida: Raphidiophryidae), the first heliozoan species with a combination of siliceous and organic skelet ...'' and '' Polyplacocystis'', have been discovered so far. References {{Taxonbar, from=Q16989009 Haptista Eukaryote families ...
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