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Canini (tribe)
Canini is a taxonomic rank which represents the dog-like tribe of the subfamily Caninae (the canines), and is sister to the fox-like tribe Vulpini. The Canini came into existence 9 million years ago. This group was first represented by ''Eucyon'', mostly by ''Eucyon davisi'' that was spread widely across North America and is basal to the other members of the tribe. Its members are informally known as true dogs. Taxonomy Members of this tribe include: Common names of most of the South American canines include "fox", based on resemblance, but they are more closely related to wolves than to ''vulpini'', the Eurasian and North American foxes. The cladogram below is based on the phylogeny of Lindblad-Toh ''et al''. (2005), modified to incorporate recent findings on ''Canis'' species, ''Lycalopex'' species, and ''Dusicyon ''Dusicyon'' is an extinct genus of South American canids. The type species is ''Dusicyon australis'', the Falkland Islands wolf. In 1914, Oldfield Thom ...
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Tortonian
The Tortonian is in the geologic time scale an age or stage of the late Miocene that spans the time between 11.608 ± 0.005 Ma and 7.246 ± 0.005 Ma (million years ago). It follows the Serravallian and is followed by the Messinian. The Tortonian roughly overlaps with the regional Pannonian Stage of the Paratethys timescale of Central Europe. It also overlaps the upper Astaracian, Vallesian and lower Turolian European land mammal ages, the upper Clarendonian and lower Hemphillian North American land mammal ages and the upper Chasicoan and lower Huayquerian South American land mammal ages. Definition The Tortonian was introduced by Swiss stratigrapher Karl Mayer-Eymar in 1858. It was named after the Italian city of Tortona in the region Piedmont. The base of the Tortonian Stage is at the last common appearance of calcareous nanoplankton '' Discoaster kugleri'' and planktonic foram '' Globigerinoides subquadratus''. It is also associated with the short normal polarized magneti ...
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Crab-eating Fox
The crab-eating fox (''Cerdocyon thous''), also known as the forest fox, wood fox, bushdog (not to be confused with the bush dog) or maikong, is an extant species of medium-sized canid endemic to the central part of South America since at least the Pleistocene epoch. Like South American foxes, which are in the genus ''Lycalopex'', it is not closely related to true foxes. ''Cerdocyon'' comes from the Greek words ''kerdo'' (meaning fox) and ''kyon'' (dog) referring to the dog- and fox-like characteristics of this animal. Taxonomy and evolution The crab-eating fox was originally described as ''Canis thous'' by Linnaeus (1766), and first placed in its current genus ''Cerdocyon'' by Hamilton-Smith in 1839. Cerdocyonina is a tribe which appeared around 6.0 million years ago (Mya) in North America as ''Ferrucyon avius'' becoming extinct by around 1.4–1.3 Mya. living about . This genus has persisted in South America from an undetermined time, possibly around 3.1 Mya, and continues ...
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Monophyly
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic groups are typically characterised by shared derived characteristics ( synapomorphies), which distinguish organisms in the clade from other organisms. An equivalent term is holophyly. The word "mono-phyly" means "one-tribe" in Greek. Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic group'' consists of all of the descendants of a common ancestor minus one or more monophyletic groups. A '' polyphyletic group'' is characterized by convergent features or habits of scientific interest (for example, night-active primates, fruit trees, aquatic insects). The features by which a polyphyletic group is differentiated from others are not inherited from a common ancestor. These definitions have taken ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Eucyon Davisi
''Eucyon'' (Greek: : good, true; : dog) is an extinct genus of medium omnivorous coyote-like canid that first appeared in the Western United States during the late Middle Miocene 10 million years ago. It was the size of a jackal and weighed around 15kg. Its species ''E. zhoui'' was one of a number of North American mammals which invaded East Asia around 5–6 million years ago, followed by the genus going extinct 3 million years ago. This genus is proposed to have given rise to genus ''Canis'' 6 million years ago. Taxonomy ''Eucyon'' was named by Tedford and Qiu in 1996. Phyletically it stood between ''Canis'' and the South American canines that would follow it. In 2009, Tedford revised its diagnosis and described two of its species, ''E. skinneri'' and ''E. davisi'', which was originally named ''Canis davisi'' by Merriam in 1911. ''Eucyon davisi'' The jackal-sized ''Eucyon'' existed in North America from 10 million YBP until the Early Pliocene. Wang and Tedford prop ...
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Vulpini
Vulpini is a taxonomic rank which represents the fox-like tribe of the subfamily Caninae (the canines), and is sister to the dog-like tribe Canini. Genera Taxonomy The taxonomy of Carnivora in general and Canidae in particular correlates with various diagnostic features of the dentition Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolo ... and basicranium. Rergarding Vulpini, Tedford has remarked: The cladogram below is based on the phylogeny of Lindblad-Toh (2005) modified to incorporate recent findings on ''Vulpes''. References Mammal tribes Canines {{Canid-stub ...
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Caninae
The Caninae, known as canines, are one of three subfamily, subfamilies found within the Canidae, canid family. The other two canid subfamilies are the extinct Borophaginae and Hesperocyoninae. The Caninae includes all living canids and their most recent fossil relatives. Their fossils were first found in North America and dated to the Oligocene era, then spreading to Asia at the end of the Miocene era, some 7 million to 8 million years ago. Taxonomy and lineage The genus ''Leptocyon'' (Greek: ''leptos'' slender + ''cyon'' dog) includes 11 species and was the first primitive canine. They were small and weighed around 2 kg. They first appeared in Sioux County, Nebraska in the Orellan era 34-32 million years ago, which was the beginning of the Oligocene. This was the same time as the appearance of the Borophaginae with whom they share features, indicating that these were two sister groups. Borophaginae skull and dentition were designed for a powerful killi ...
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Tribe (biology)
In biology, a tribe is a taxonomic rank above genus, but below family and subfamily. It is sometimes subdivided into subtribes. By convention, all taxonomic ranks from genus upwards are capitalized, including both tribe and subtribe. In zoology, the standard ending for the name of a zoological tribe is "-ini". Examples include the tribes Caprini (goat-antelopes), Hominini (hominins), Bombini (bumblebees), and Thunnini (tunas). The tribe Hominini is divided into subtribes by some scientists; subtribe Hominina then comprises "humans". The standard ending for the name of a zoological subtribe is "-ina". In botany, the standard ending for the name of a botanical tribe is "-eae". Examples include the tribes Acalypheae and Hyacintheae. The tribe Hyacintheae is divided into subtribes, including the subtribe Massoniinae. The standard ending for the name of a botanical subtribe is "-inae". In bacteriology, the form of tribe names is as in botany, e.g., Pseudomonadeae, based on the ge ...
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Taxonomic Rank
In biological classification, taxonomic rank is the relative level of a group of organisms (a taxon) in an ancestral or hereditary hierarchy. A common system consists of species, genus, family (biology), family, order (biology), order, class (biology), class, phylum (biology), phylum, kingdom (biology), kingdom, domain (biology), domain. While older approaches to taxonomic classification were phenomenological, forming groups on the basis of similarities in appearance, organic structure and behaviour, methods based on genetic analysis have opened the road to cladistics. A given rank subsumes under it less general categories, that is, more specific descriptions of life forms. Above it, each rank is classified within more general categories of organisms and groups of organisms related to each other through inheritance of phenotypic trait, traits or features from common ancestors. The rank of any ''species'' and the description of its ''genus'' is ''basic''; which means that to iden ...
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Theriodictis
''Theriodictis'' is an extinct genus of hypercarnivorous wolf-like canid endemic to South America during the Pleistocene, living from 1.2 Ma- 500,000 years ago and existing for approximately . Fossil distribution The fossil remains are confined to the Tarija Formation of Bolivia, the Chui Formation of southern Brazil, and the Yupoí Formation of northern Argentina.''Theriodictis''
at .org
The species ''T. tarijense'' was transferred to the genus '''' upon phylogenetic analysis.


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Protocyon
''Protocyon'' is an extinct genus of large canid endemic to South and North America during the Late Pleistocene living from 781 to 12thousand years ago. Description ''Protocyon'' was a hypercarnivore, suggested by its dental adaptations. Like many other large canids, it was most likely a pack hunter. It hunted the medium-sized grazers and browsers, and bite marks on fossils suggest that it may have hunted '' Glyptotherium''. The find of a molar tooth found in Santa Vitória do Palmar in Brazil suggests a weight of between for this particular specimen. , modest in size compared to other canids including the dire wolf, despite its modest size isotopic analysis shows a dietary overlap with Smilodon populator, which implies it actively hunted much larger prey animals since S. populator weighed at least 400kg. Taxonomy ''Protocyon'' was named by Giebel in 1855 and assigned to Canidae by Carroll in 1988. A member of the Cerdocyonina lineage, its closest living relative might be the bus ...
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Dusicyon
''Dusicyon'' is an extinct genus of South American canids. The type species is ''Dusicyon australis'', the Falkland Islands wolf. In 1914, Oldfield Thomas established this genus, in which he included the culpeo and other South American foxes. These other canids were removed to '' Lycalopex'' by Langguth in 1975. '' Dusicyon avus'', widely distributed in the late Pleistocene from Uruguay through Buenos Aires Province to southernmost Chile, is the closest known relative of the Falkland Islands wolf; the two lineages split only about 16,000 years ago. It died out in the late Holocene, earlier estimates suggested about 2,980 years ago on the island of Tierra del Fuego and almost 1,700 years ago in the continent. More recent research confirms much later extinction dates, with the latest confirmed records in the Pampean Region being 700 BP (1232–1397 AD) and southernmost Patagonia at 400 years BP (1454–1626 AD). There is still much debate about the classification of ''"Dusicyo ...
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