''Protocyon'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

of large canid endemic to

South South is one of the cardinal directions or compass points. The direction is the opposite of north and is perpendicular to both east and west. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Proto-Germanic ''*sunþa ...

and

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...

during the

Late Pleistocene The Late Pleistocene is an unofficial age in the international geologic timescale in chronostratigraphy, also known as Upper Pleistocene from a stratigraphic perspective. It is intended to be the fourth division of the Pleistocene Epoch withi ...

living from 781 to 12thousand years ago.

Description

''Protocyon'' was a

hypercarnivore A hypercarnivore is an animal which has a diet that is more than 70% meat, either via active predation or by scavenging. The remaining non-meat diet may consist of non-animal foods such as fungi, fruits or other plant material. Some extant exam ...

, suggested by its dental adaptations. Like many other large canids, it was most likely a pack hunter. It hunted the medium-sized grazers and browsers, and bite marks on fossils suggest that it may have hunted ''

Glyptotherium ''Glyptotherium'' (from Greek for 'grooved or carved beast') is a genus of glyptodont (an extinct group of large, herbivorous armadillos) that lived from the Early Pliocene, about 4.9 million years ago, to the Early Holocene, around 7,000 yea ...

''. The find of a molar tooth found in

Santa Vitória do Palmar Santa Vitória do Palmar (lit. "Saint Victoria of the Palms") is a Brazilian city and municipality. It is the southernmost municipality in Brazil, located in the state of Rio Grande do Sul. To the west of the municipality is the Lagoa Mirim and ...

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

suggests a weight of between for this particular specimen. , modest in size compared to other canids including the dire wolf, despite its modest size isotopic analysis shows a dietary overlap with Smilodon populator, which implies it actively hunted much larger prey animals since S. populator weighed at least 400kg.

Taxonomy

''Protocyon'' was named by Giebel in 1855 and assigned to Canidae by Carroll in 1988. A member of the

Cerdocyonina Cerdocyonina is an extant subtribe of the canines and is exclusively endemic to the Americas. Often described to be "fox-like" in appearance and behavior, they are more closely related to the wolf-like canids such as ''Canis'' than they are to ...

lineage, its closest living relative might be the

bush dog The bush dog (''Speothos venaticus'') is a canine found in Central and South America. In spite of its extensive range, it is very rare in most areas except in Suriname, Guyana and Peru; it was first identified by Peter Wilhelm Lund from foss ...

Fossil distribution

Fossils of ''Protocyon'' have been found in the Ñuapua and Tarija Formations of

Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...

, the Vorohue Formation of

Buenos Aires Buenos Aires ( or ; ), officially the Autonomous City of Buenos Aires ( es, link=no, Ciudad Autónoma de Buenos Aires), is the capital and primate city of Argentina. The city is located on the western shore of the Río de la Plata, on South ...

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the List of South American countries by area, second-largest ...

, Santa Elena Peninsula of

Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ' ...

, Sopas Formation of

Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...

Mene de Inciarte Tar Seep The moonfish of the genus ''Mene'', the sole extant genus of the family Menidae, are disk-shaped fish which bear a vague resemblance to gourami, thanks to their thread-like pelvic fins. Today, the genus is represented only by ''Mene maculata'' ...

Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...

and various sites in

, among others the Jandaíra Formation. Canid fossil material from the Hoyo Negro pit in the Sac Actun cave system (

Mexico Mexico (Spanish language, Spanish: México), officially the United Mexican States, is a List of sovereign states, country in the southern portion of North America. It is borders of Mexico, bordered to the north by the United States; to the so ...

), initially identified as remains of a

coyote The coyote (''Canis latrans'') is a species of canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecological nich ...

, was reinterpreted as remains of ''Protocyon troglodytes'' by ', indicating that this taxon was also present in the southern part of North America.

References

{{Taxonbar, from=Q142407 Cerdocyonina Prehistoric canines Prehistoric carnivoran genera Pleistocene carnivorans Pleistocene mammals of South America Pleistocene mammals of North America Lujanian Pleistocene Argentina Fossils of Argentina Pleistocene Bolivia Fossils of Bolivia Pleistocene Brazil Fossils of Brazil Pleistocene Ecuador Fossils of Ecuador Pleistocene Mexico Fossils of Mexico Pleistocene Uruguay Fossils of Uruguay Pleistocene Venezuela Fossils of Venezuela Fossil taxa described in 1855