Cambridge Greensand Formation
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Cambridge Greensand Formation
The Cambridge Greensand is a geological unit in England whose strata are earliest Cenomanian in age. It lies above the erosive contact between the Gault Formation and the Chalk Group in the vicinity of Cambridgeshire, and technically forms the lowest member bed of the West Melbury Marly Chalk Formation.Cambridge Greensand
at BGS
It is a remanié deposit, containing reworked fossils of late age, including those of dinosaurs and pterosaurs.


Description

The lithology is made out of

Stratigraphic Unit
A stratigraphic unit is a volume of rock of identifiable origin and relative age range that is defined by the distinctive and dominant, easily mapped and recognizable petrographic, lithologic or paleontologic features (facies) that characterize it. Units must be ''mappable'' and ''distinct'' from one another, but the contact need not be particularly distinct. For instance, a unit may be defined by terms such as "when the sandstone component exceeds 75%". Lithostratigraphic units Sequences of sedimentary and volcanic rocks are subdivided the basis of their shared or associated lithology. Formally identified lithostratigraphic units are structured in a hierarchy of lithostratigraphic rank, higher rank units generally comprising two or more units of lower rank. Going from smaller to larger in rank, the main lithostratigraphic ranks are Bed, Member, Formation, Group and Supergroup. Formal names of lithostratigraphic units are assigned by geological surveys. Units of formation or hi ...
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Foraminifera
Foraminifera (; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular Ectoplasm (cell biology), ectoplasm for catching food and other uses; and commonly an external shell (called a "Test (biology), test") of diverse forms and materials. Tests of chitin (found in some simple genera, and Textularia in particular) are believed to be the most primitive type. Most foraminifera are marine, the majority of which live on or within the seafloor sediment (i.e., are benthos, benthic), while a smaller number float in the water column at various depths (i.e., are planktonic), which belong to the suborder Globigerinina. Fewer are known from freshwater or brackish conditions, and some very few (nonaquatic) soil species have been identified through molecular analysis of small subunit ribosomal DNA. Foraminifera typically produce a test (biology), test, or shell, which can have eithe ...
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Lonchodraco
''Lonchodraco'' is a genus of lonchodraconid Pterodactyloidea, pterodactyloid pterosaur from the Late Cretaceous of southern England. The genus includes species that were previously assigned to other genera. Discovery and naming In 1846, James Scott Bowerbank named and described some remains found in a chalk pit at Burham near Maidstone in Kent, as a new species of ''Pterodactylus'': ''Pterodactylus giganteus''. The specific name (zoology), specific name means "the gigantic one" in Latin. The same pit generated remains of ''Cimoliopterus, Pterodactylus cuvieri''. In 1848, Bowerbank published a histological study of the bone structure of ''P. giganteus''. At the time, the British Association Code of 1843 allowed to change names if they were inappropriate. In 1850, Richard Owen, considering the species not to have been particularly large, and renamed it into ''Pterodactylus conirostris''; the specific name meaning "cone-snouted", which was based on the conical snout of specimen ...
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Draigwenia
''Draigwenia'' (meaning "white dragon" from the Welsh ''ddraig wen'', referring to a dragon associated with the Anglo-Saxons) is an extinct genus of pterosaur known from a jaw fragment found in the Late Cretaceous Cambridge Greensand in the United Kingdom. The fossil was likely reworked from an Early Cretaceous layer that can be dated to the Albian. It currently contains a single species, ''Draigwenia platystomus''. History The holotype CAMSM B54835, was described in 1870 by Harry Govier Seeley as the holotype of his species ''Ornithocheirus platystomus''. In 1914 Reginald Walter Hooley moved ''O''. ''platystomus'' to ''Amblydectes''. It was assigned by David Unwin in 2001 to the genus ''Lonchodectes'',Unwin D.M., (2001), "An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England", ''Mitteilungen aus dem Museum für Naturkunde in Berlin'', Geowissenschaftliche Reihe 4: 189–221 and in 2013 was excluded from the Lonchodraconidae. In ...
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Camposipterus Sedgwickii
''Aerodraco'' (meaning "air dragon") is a genus of anhanguerid pterosaur from the Albian–Cenomanian-age Cambridge Greensand of England. It contains only one species, ''Aerodraco sedgwickii''. It was originally assigned to the genus ''Pterodactylus''. Discovery and naming In 1859, Sir Richard Owen named pterosaur material from the Cambridge Greensand of England as ''Pterodactylus sedgwickii''.R. Owen. 1859. Supplement (No. I) to the Monograph on the Fossil Reptilia of the Cretaceous Formations. Palaeontographical Society Monographs 1–19 At the time, ''Pterodactylus'' was a wastebasket taxon; all sorts of unrelated pterosaurs were assigned to that genus. In 1870, Harry Seeley reassigned it to ''Ornithocheirus'', another wastebasket taxon. Its specific name honors Adam Sedgwick. It was in 1869 renamed by Seeley into a ''Ptenodactylus sedgwickii'',Seeley, H.G., 1869, ''Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arrange ...
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Camposipterus
''Camposipterus'' is a genus of pterodactyloid pterosaur from the Early Cretaceous of England. Fossil remains of ''Camposipterus'' dated back to the Early Cretaceous, about 112 million years ago. Discovery and naming In 1869, Harry Govier Seeley, based on a fossil found at Haslingfield, Cambridgeshire, named ''Ptenodactylus nasutus'',Seeley, H.G., 1869, ''Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arranged in the Woodwardian Museum of the University of Cambridge''. Deighton, Bell and Co., Cambridge, xxiii + 143 pp at the same time disclaiming the name which makes it invalid by modern standards. In 1870, Seeley had realized that the generic name ''Ptenodactylus'' had been preoccupied, so he renamed the species into ''Ornithocheirus nasutus''.Seeley, H.G., 1870, ''The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwar ...
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Camposipterus Colorhinus
''Camposipterus'' is a genus of pterodactyloid pterosaur from the Early Cretaceous of England. Fossil remains of ''Camposipterus'' dated back to the Early Cretaceous, about 112 million years ago. Discovery and naming In 1869, Harry Govier Seeley, based on a fossil found at Haslingfield, Cambridgeshire, named ''Ptenodactylus nasutus'',Seeley, H.G., 1869, ''Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arranged in the Woodwardian Museum of the University of Cambridge''. Deighton, Bell and Co., Cambridge, xxiii + 143 pp at the same time disclaiming the name which makes it invalid by modern standards. In 1870, Seeley had realized that the generic name ''Ptenodactylus'' had been preoccupied, so he renamed the species into ''Ornithocheirus nasutus''.Seeley, H.G., 1870, ''The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwar ...
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Amblydectes
''Amblydectes'' is a genus of pterosaur known from jaw fragments. It apparently had a jaw flattened towards the tip and triangular in cross-section. It has at times been synonymized with ''Coloborhynchus'', ''Criorhynchus'', ''Lonchodectes'', or ''Ornithocheirus''. A 2013 study found ''A. crassidens'' and ''A. eurygnathus'' to be ''nomina dubia'', with ''A. platystomus'' possibly belonging to a separate, yet unnamed genus. A 2021 study found ''A. crassidens'' to be a valid genus within Anhangueridae, while ''A. platystomus'' was placed in the new genus ''Draigwenia''. ''A. eurygnathus'' was found to possibly be a junior synonym of ''A. crassidens''. See also * Timeline of pterosaur research This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic era. Although pterosaurs w ... References Pterodactyloids Early ...
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Amblydectes Crassidens
''Amblydectes'' is a genus of pterosaur known from jaw fragments. It apparently had a jaw flattened towards the tip and triangular in cross-section. It has at times been synonymized with ''Coloborhynchus'', ''Criorhynchus'', ''Lonchodectes'', or ''Ornithocheirus''. A 2013 study found ''A. crassidens'' and ''A. eurygnathus'' to be ''nomina dubia'', with ''A. platystomus'' possibly belonging to a separate, yet unnamed genus. A 2021 study found ''A. crassidens'' to be a valid genus within Anhangueridae, while ''A. platystomus'' was placed in the new genus ''Draigwenia''. ''A. eurygnathus'' was found to possibly be a junior synonym of ''A. crassidens''. See also * Timeline of pterosaur research This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic era. Although pterosaurs w ... References Pterodactyloids Early ...
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Macrurosaurus
''Macrurosaurus'' (meaning "large-tailed lizard") is the name given to a genus of dinosaur from the Early Cretaceous. It was a titanosauriform which lived in what is now England. The type species, ''M. semnus'', was named in 1876. A second species, ''M. platypus'', may also exist. History The genus ''Macrurosaurus'' was named by Harry Govier Seeley in 1869 in his index of fossils from the Cambridge Greensand. In 1876 Seeley more thoroughly described the type species, ''Macrurosaurus semnus'', making the name valid.H.G. Seeley, 1876, "On ''Macrurosaurus semnus'' (Seeley), a long tailed animal with procoelous vertebrae from the Cambridge Upper Greensand, preserved in the Woodwardian Museum of the University of Cambridge", ''Quarterly Journal of the Geological Society of London'' 32: 440-444 The generic name is derived from Greek ''makros'', "large", and ''oura'', "tail". The specific name is derived from Greek ''semnos'', "stately" or "impressive". A second species, ''M. platypus ...
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Trachodon Cantabrigiensis
This list of informally named dinosaurs is a listing of dinosaurs (excluding Aves; birds and their extinct relatives) that have never been given formally published scientific names. This list only includes names that were not properly published (" unavailable names") and have not since been published under a valid name (see list of dinosaur genera for valid names). The following types of names are present on this list: * ''Nomen nudum'', Latin for "naked name": A name that has appeared in print but has not yet been formally published by the standards of the International Commission on Zoological Nomenclature. ''Nomina nuda'' (the plural form) are invalid, and are therefore not italicized as a proper generic name would be. * '' Nomen manuscriptum'', Latin for "manuscript name": A name that appears in manuscript but was not formally published. A ''nomen manuscriptum'' is equivalent to a ''nomen nudum'' for everything except the method of publication, and description. * '' Nomen ex ...
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Eucercosaurus
''Eucercosaurus'' (meaning "good-tailed lizard") is the name given to a genus of ornithopod dinosaur from the Albian stage of the Early Cretaceous. It was an ornithopod discovered in the Cambridge Greensand of England and is known from 19 centra, 3 sacrals, 4 dorsals and 12 caudals, and a neural arch found near Trumpington, Cambridgeshire. The type species, ''E. tanyspondylus'', was described by British paleontologist Harry Seeley in 1879. It is considered a dubious name, and was once considered an ankylosaur. According to a 2020 study, ''Eucercosaurus'' and ''Syngonosaurus'' were basal iguanodontia Iguanodontia (the iguanodonts) is a clade of herbivorous dinosaurs that lived from the Middle Jurassic to Late Cretaceous. Some members include ''Camptosaurus'', ''Dryosaurus'', ''Iguanodon'', ''Tenontosaurus'', and the hadrosaurids or "duck-bi ...ns. References Ornithopods Albian life Early Cretaceous dinosaurs of Europe Cretaceous England Fossils of England Fossil ta ...
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