Amblydectes
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Amblydectes
''Amblydectes'' is a genus of pterosaur known from jaw fragments. It apparently had a jaw flattened towards the tip and triangular in cross-section. It has at times been synonymized with ''Coloborhynchus'', ''Criorhynchus'', ''Lonchodectes'', or ''Ornithocheirus''. A 2013 study found ''A. crassidens'' and ''A. eurygnathus'' to be ''nomina dubia'', with ''A. platystomus'' possibly belonging to a separate, yet unnamed genus. A 2021 study found ''A. crassidens'' to be a valid genus within Anhangueridae, while ''A. platystomus'' was placed in the new genus ''Draigwenia''. ''A. eurygnathus'' was found to possibly be a junior synonym of ''A. crassidens''. See also * Timeline of pterosaur research This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic era. Although pterosaurs w ... References Pterodactyloids Early ...
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Amblydectes Eurygnathus
''Amblydectes'' is a genus of pterosaur known from jaw fragments. It apparently had a jaw flattened towards the tip and triangular in cross-section. It has at times been synonymized with ''Coloborhynchus'', ''Criorhynchus'', ''Lonchodectes'', or ''Ornithocheirus''. A 2013 study found ''A. crassidens'' and ''A. eurygnathus'' to be ''nomina dubia'', with ''A. platystomus'' possibly belonging to a separate, yet unnamed genus. A 2021 study found ''A. crassidens'' to be a valid genus within Anhangueridae, while ''A. platystomus'' was placed in the new genus ''Draigwenia''. ''A. eurygnathus'' was found to possibly be a junior synonym of ''A. crassidens''. See also * Timeline of pterosaur research References

Pterodactyloids Early Cretaceous pterosaurs of Europe Taxa named by Reginald Hooley Fossil taxa described in 1914 {{pterosaur-stub ...
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Draigwenia
''Draigwenia'' (meaning "white dragon" from the Welsh ''ddraig wen'', referring to a dragon associated with the Anglo-Saxons) is an extinct genus of pterosaur known from a jaw fragment found in the Late Cretaceous Cambridge Greensand in the United Kingdom. The fossil was likely reworked from an Early Cretaceous layer that can be dated to the Albian. It currently contains a single species, ''Draigwenia platystomus''. History The holotype CAMSM B54835, was described in 1870 by Harry Govier Seeley as the holotype of his species ''Ornithocheirus platystomus''. In 1914 Reginald Walter Hooley moved ''O''. ''platystomus'' to ''Amblydectes''. It was assigned by David Unwin in 2001 to the genus ''Lonchodectes'',Unwin D.M., (2001), "An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England", ''Mitteilungen aus dem Museum für Naturkunde in Berlin'', Geowissenschaftliche Reihe 4: 189–221 and in 2013 was excluded from the Lonchodraconidae. In ...
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Ornithocheirus
''Ornithocheirus'' (from Ancient Greek "ὄρνις", meaning ''bird'', and "χεῖρ", meaning ''hand'') is a pterosaur genus known from fragmentary fossil remains uncovered from sediments in the UK and possibly Morocco. Several species have been referred to the genus, most of which are now considered as dubious species, or members of different genera, and the genus is now often considered to include only the type species, ''Ornithocheirus simus''. Species have been referred to ''Ornithocheirus'' from the mid-Cretaceous period of both Europe and South America, but ''O. simus'' is known only from the UK, though a specimen referred to ''O.'' cf. ''simus'' is also known from Morocco. Because ''O. simus'' was originally named based on poorly preserved fossil material, the genus ''Ornithocheirus'' has suffered enduring problems of zoological nomenclature. Fossil remains of ''Ornithocheirus'' have been recovered mainly from the Cambridge Greensand of England, dating to the beginni ...
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Criorhynchus
''Ornithocheirus'' (from Ancient Greek "ὄρνις", meaning ''bird'', and "χεῖρ", meaning ''hand'') is a pterosaur genus known from fragmentary fossil remains uncovered from sediments in the UK and possibly Morocco. Several species have been referred to the genus, most of which are now considered as dubious species, or members of different genera, and the genus is now often considered to include only the type species, ''Ornithocheirus simus''. Species have been referred to ''Ornithocheirus'' from the mid-Cretaceous period of both Europe and South America, but ''O. simus'' is known only from the UK, though a specimen referred to ''O.'' cf. ''simus'' is also known from Morocco. Because ''O. simus'' was originally named based on poorly preserved fossil material, the genus ''Ornithocheirus'' has suffered enduring problems of zoological nomenclature. Fossil remains of ''Ornithocheirus'' have been recovered mainly from the Cambridge Greensand of England, dating to the beginni ...
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Lonchodectes
''Lonchodectes'' (meaning "lance biter") was a genus of lonchodectid pterosaur from several formations dating to the Turonian (Late Cretaceous) of England, mostly in the area around Kent. The species belonging to it had been assigned to ''Ornithocheirus'' until David Unwin's work of the 1990s and 2000s.Kellner, A.W.A. (2003). Pterosaur phylogeny and comments on the evolutionary history of the group: In: Buffetaut, E., and Mazin, J.-M. (Eds.). ''Evolution and Palaeobiology of Pterosaurs''. Geological Society Special Publication 217:105-137. 1-86239-143-2. Several potential species are known; most are based on scrappy remains, and have gone through several other generic assignments. The genus is part of the complex taxonomy issues surrounding Early Cretaceous pterosaurs from Brazil and England, such as ''Amblydectes'', ''Anhanguera'', ''Coloborhynchus'', and ''Ornithocheirus''. History and species Numerous species have been referred to this genus over time, and only those more ...
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Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin ''creta'', "chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation ''Kreide''. The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now- extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was ice free, and forests extended to the poles. During this time, new groups of mammals and birds appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth b ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Coloborhynchus
''Coloborhynchus'' is a genus of pterodactyloid pterosaur belonging to the family Anhangueridae, though it has also been recovered as a member of the Ornithocheiridae in some studies. ''Coloborhynchus'' is known from the Lower Cretaceous of England (Valanginian age, 140 to 136 million years ago), and depending on which species are included, possibly the Albian and Cenomanian ages (113 to 93.9 million years ago) as well. ''Coloborhynchus'' was once thought to be the largest known toothed pterosaur, however, a specimen of the closely related ''Tropeognathus'' is now thought to have had a larger wingspan. History and classification Like many ornithocheiroid pterosaurs named during the 19th century, ''Coloborhynchus'' has a highly convoluted history of classification. Over the years numerous species have been assigned to it, and often, species have been shuffled between ''Coloborhynchus'' and related genera by various researchers. In 1874 Richard Owen, rejecting the creation by H ...
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Nomina Dubia
In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a specimen belongs to that group or not. This may happen if the original type series (i. e. holotype, isotype, syntype or paratype) is lost or destroyed. The zoological and botanical codes allow for a new type specimen, or neotype, to be chosen in this case. A name may also be considered a ''nomen dubium'' if its name-bearing type is fragmentary or lacking important diagnostic features (this is often the case for species known only as fossils). To preserve stability of names, the ''International Code of Zoological Nomenclature'' allows a new type specimen, or neotype, to be chosen for a ''nomen dubium'' in this case. 75.5. Replacement of unidentifiable name-bearing type by a neotype. When an author considers that the taxonomic identity of a ...
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Anhangueridae
Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a lateral expansion in the distal rostrum. The same study presented a cladistic analysis, for which an "agreement subtree" was calculated. The Anhangueridae was found to be sister taxon to the large crested ''Tropeognathus''. Relationships There are competing theories of ornithocheiromorph phylogeny (evolutionary relationships). Below is cladogram following a topology recovered by Brian Andres, using the most recent iteration of his data set. The cladogram below follows Pêgas ''et al.'' (2019), who recovered Anhangueridae as a much more inclusive group. The analysis found most of the ornithocheirids falling into this family, while ''Ornithocheirus'' itself was recovered as a basal member of Ornithocheirae Ornithocheirae is an extinct clad ...
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Junior Synonym
The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linnaeus was the first to give a scientific name (under the currently used system of scientific nomenclature) to the Norway spruce, which he called ''Pinus abies''. This name is no longer in use, so it is now a synonym of the current scientific name, ''Picea abies''. * In zoology, moving a species from one genus to another results in a different binomen, but the name is considered an alternative combination rather than a synonym. The concept of synonymy in zoology is reserved for two names at the same rank that refers to a taxon at that rank - for example, the name ''Papilio prorsa'' Linnaeus, 1758 is a junior synonym of ''Papilio levana'' Linnaeus, 1758, being names for different seasonal forms of the species now referred to as ''Araschnia lev ...
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