Burgess Shale Type Fauna
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Burgess Shale Type Fauna
A number of assemblages bear fossil assemblages similar in character to that of the Burgess Shale. While many are also preserved in a similar fashion to the Burgess Shale, the term "Burgess Shale-type fauna" covers assemblages based on taxonomic criteria only. Extent The fauna of the middle Cambrian has a cosmopolitan range. All assemblages preserving soft-part anatomy have a very similar fauna, even though they span almost every continent. The wide distribution has been attributed to the advent of pelagic larvae. Composition The fauna is composed of a range of soft-bodied organisms; creatures with hard, mineralised skeletons are rare, although trilobites are quite commonly found. The major soft-bodied groups are sponges, palaeoscolecid worms, lobopods, arthropods and anomalocaridids. Assemblages are typically diverse, with the most famous localities each containing in the region of 150 described species. The fauna of the Burgess Shale lived in the photic zone, as bottom-dwelling ...
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Burgess Shale
The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At old (middle Cambrian), it is one of the earliest fossil beds containing soft-part imprints. The rock unit is a black shale and crops out at a number of localities near the town of Field in Yoho National Park Yoho National Park ( ) is a National Parks of Canada, national park of Canada. It is located within the Canadian Rockies, Rocky Mountains along the western slope of the Continental Divide of the Americas in southeastern British Columbia, bordered ... and the Kicking Horse Pass. Another outcrop is in Kootenay National Park 42 km to the south. History and significance The Burgess Shale was discovered by palaeontologist Charles Doolittle Walcott, Charles Walcott on 30 August 1909, towards the end of the season's fieldwork. He returned in 1910 with his sons, daughter, and wif ...
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Pambdelurion Whittingtoni
''Pambdelurion'' is an extinct genus of panarthropod from the Cambrian aged Sirius Passet site in northern Greenland. Like the morphologically similar ''Kerygmachela'' from the same locality, ''Pambdelurion'' is thought to be closely related to arthropods, combining characteristics of " lobopodians" with those of primitive arthropods. Description ''Pambdelurion'' was large for a Cambrian animal, and is estimated to have reached a length of . ''Omnidens'', an organism from China that closely resembles ''Pambdelurion'' and may even be synonymous with it, reached even larger sizes, estimated to be based on the proportions of ''Pambdelurion''. The head of ''Pambdelurion'' bore a large pair of frontal appendages, homologous to the antennae of onychophorans and frontal appendages of radiodonts. These frontal appendages were weakly muscled and relatively soft, suggesting they may have served primarily as sensory organs, rather than for grasping prey. Between the appendages are a pair ...
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Yunnanozoon
''Yunnanozoon lividum'' (Yunnan + Greek ζῷον ''zôion'', ''lividum''; "livid animal of Yunnan") is an extinct species of possible vertebrate or chordate from the Lower Cambrian, Chengjiang biota of Yunnan province, China. It is thought of as a deuterostome suspected of being either a hemichordate or chordate. In 2022, a study reanalyzed fossils of ''Yunnanozoon'' and found it to be one of the earliest members of the vertebrate family tree. ''Yunnanozoon'' is similar to the form ''Haikouella'', which is almost certainly a chordate. Still, there are anatomical differences from ''Haikouella'', including a smaller stomach and much larger (1 mm) pharyngeal teeth. It is by no means certain whether Yunannozoon possessed features such as a heart, gills, etc., which are seen in well-preserved specimens of ''Haikouella''. ''Yunnanozoon'' somewhat resembles the Middle Cambrian ''Pikaia'' from the Burgess shale of British Columbia in Canada. Thirteen pairs of symmetrically arra ...
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Lamprey
Lampreys (sometimes inaccurately called lamprey eels) are an ancient extant lineage of jawless fish of the order Petromyzontiformes , placed in the superclass Cyclostomata. The adult lamprey may be characterized by a toothed, funnel-like sucking mouth. The common name "lamprey" is probably derived from Latin , which may mean "stone licker" ( "to lick" + "stone"), though the etymology is uncertain. ''Lamprey'' is sometimes seen for the plural form. There are about 38 known extant species of lampreys and five known extinct species. Parasitic carnivorous species are the most well-known, and feed by boring into the flesh of other fish to suck their blood; but only 18 species of lampreys engage in this micropredatory lifestyle. Of the 18 carnivorous species, nine migrate from saltwater to freshwater to breed (some of them also have freshwater populations), and nine live exclusively in freshwater. All non-carnivorous forms are freshwater species. Adults of the non-carnivorous ...
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Haikouichthys
''Haikouichthys'' is an extinct genus of craniate (animals with notochords and distinct heads) that lived 518 million years ago, during the Cambrian explosion of multicellular life. ''Haikouichthys'' had a defined skull and other characteristics that have led paleontologists to label it a true craniate, and even to be popularly characterized as one of the Prehistoric fish, earliest fishes. Cladistics, Cladistic analysis indicates that the animal is probably a basal chordate or a basal craniate; but it does not possess sufficient features to be included uncontroversially even in either stem group. It was formally described in 1999. Description ''Haikouichthys'' is about long and is narrower than ''Myllokunmingia'', another Wiktionary:putative#English, putative chordate that comes from the same beds. The holotype of ''Haikouichthys ercaicunensis'' was found in the Yuansshan member of the Maotianshan Shales, Qiongzhusi Formation in the 'Eoredlichia' Zone near Haikou at Ercaicun, ...
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Agnathid
Agnatha (, Ancient Greek 'without jaws') is an infraphylum of jawless fish in the phylum Chordata, subphylum Vertebrata, consisting of both present (cyclostomes) and extinct (conodonts and ostracoderms) species. Among recent animals, cyclostomes are sister to all vertebrates with jaws, known as gnathostomes. Recent molecular data, both from rRNA and from mtDNA as well as embryological data, strongly supports the hypothesis that living agnathans, the cyclostomes, are monophyletic. The oldest fossil agnathans appeared in the Cambrian, and two groups still survive today: the lampreys and the hagfish, comprising about 120 species in total. Hagfish are considered members of the subphylum Vertebrata, because they secondarily lost vertebrae; before this event was inferred from molecular and developmental data, the group Craniata was created by Linnaeus (and is still sometimes used as a strictly morphological descriptor) to reference hagfish plus vertebrates. While a few scientist ...
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Myllokunmingia
''Myllokunmingia'' is a genus of basal chordate from the Lower Cambrian Maotianshan shales of China 518 to 490 mya and is thought to be a vertebrate, although this is not conclusively proven. The species M. fengjiaoa is 28 mm long and 6 mm high. It is among the oldest possible craniates, found in the lower Cambrian Chengjiang (). It appears to have a skull and skeletal structures made of cartilage. There is no sign of biomineralization of the skeletal elements. The holotype was found in the Yuanshan member of the Qiongzhusi Formation in the Eoredlichia Zone near Haikou at Ercaicun, Kunming City, Yunnan, China. Only one species is known – ''Myllokunmingia fengjiaoa'' (Shu, Zhang & Han). Related creatures are ''Haikouichthys'' and '' Zhongjianichthys''. The animal has a distinct head and trunk with a forward sail-like (1.5 mm) dorsal fin and a ventral finfold (probably paired) further back. The maximum height of M. fengijaoa is at 6mm. The maximum height point i ...
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Chordata
A chordate () is an animal of the phylum Chordata (). All chordates possess, at some point during their larval or adult stages, five synapomorphies, or primary physical characteristics, that distinguish them from all the other taxa. These five synapomorphies include a notochord, dorsal hollow nerve cord, endostyle or thyroid, pharyngeal slits, and a post-anal tail. The name “chordate” comes from the first of these synapomorphies, the notochord, which plays a significant role in chordate structure and movement. Chordates are also Bilateral symmetry, bilaterally symmetric, have a coelom, possess a circulatory system, and exhibit Metameric, metameric segmentation. In addition to the morphological characteristics used to define chordates, analysis of genome sequences has identified two conserved signature indels (CSIs) in their proteins: cyclophilin-like protein and mitochondrial inner membrane protease ATP23, which are exclusively shared by all vertebrates, tunicates and cep ...
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Maotianshan Shale
The Maotianshan Shales are a series of Early Cambrian deposits in the Chiungchussu Formation, famous for their '' Konservat Lagerstätten'', deposits known for the exceptional preservation of fossilized organisms or traces. The Maotianshan Shales form one of some forty Cambrian fossil locations worldwide exhibiting exquisite preservation of rarely preserved, non-mineralized soft tissue, comparable to the fossils of the Burgess Shale. They take their name from Maotianshan Hill (, Literal meaning: Hat Sky Mountain) in Chengjiang County, Yunnan Province, China. The most famous assemblage of organisms are referred to as the Chengjiang biota for the multiple scattered fossil sites in Chengjiang. The age of the Chengjiang Lagerstätte is locally termed Qiongzhusian, a stage correlated to the late Atdabanian Stage in Siberian sequences of the middle of the Early Cambrian. The shales date to ≤. The shales also contain the slightly younger Guanshan biota from Malong District in Yunnan ...
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Yunnan
Yunnan , () is a landlocked Provinces of China, province in Southwest China, the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The province borders the Chinese provinces of Guizhou, Sichuan, autonomous regions of Guangxi, and Tibet Autonomous Region, Tibet as well as Southeast Asian countries: Vietnam, Laos, and Myanmar. Yunnan is China's fourth least developed province based on disposable income per capita in 2014. Yunnan is situated in a mountainous area, with high elevations in the northwest and low elevations in the southeast. Most of the population lives in the eastern part of the province. In the west, the altitude can vary from the mountain peaks to river valleys by as much as . Yunnan is rich in natural resources and has the largest diversity of plant life in China. Of the approximately 30,000 species of Vascular plant, higher plants in China, Yu ...
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Chengjiang Fauna
The Maotianshan Shales are a series of Early Cambrian deposits in the Chiungchussu Formation, famous for their '' Konservat Lagerstätten'', deposits known for the exceptional preservation of fossilized organisms or traces. The Maotianshan Shales form one of some forty Cambrian fossil locations worldwide exhibiting exquisite preservation of rarely preserved, non-mineralized soft tissue, comparable to the fossils of the Burgess Shale. They take their name from Maotianshan Hill (, Literal meaning: Hat Sky Mountain) in Chengjiang County, Yunnan Province, China. The most famous assemblage of organisms are referred to as the Chengjiang biota for the multiple scattered fossil sites in Chengjiang. The age of the Chengjiang Lagerstätte is locally termed Qiongzhusian, a stage correlated to the late Atdabanian Stage in Siberian sequences of the middle of the Early Cambrian. The shales date to ≤. The shales also contain the slightly younger Guanshan biota from Malong District in Yunnan ...
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Biramous
The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments (called podomeres) are of Latin origin, and may be confused with terms for bones: ''coxa'' (meaning hip, plural ''coxae''), ''trochanter'', ''femur'' (plural ''femora''), ''tibia'' (plural ''tibiae''), ''tarsus'' (plural ''tarsi''), ''ischium'' (plural ''ischia''), ''metatarsus'', ''carpus'', ''dactylus'' (meaning finger), ''patella'' (plural ''patellae''). Homologies of leg segments between groups are difficult to prove and are the source of much argument. Some authors posit up to eleven segments per leg for the most recent common ancestor of extant arthropods but modern arthropods have eight or fewer. It has been argued that the ancestral leg need not have been so complex, and that other events, such as successive loss of function of a ''Hox''-gene, could result in parallel gains of leg segments. In arthropods, each of the leg segments artic ...
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