Myllokunmingia
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Myllokunmingia
''Myllokunmingia'' is a genus of basal chordate from the Lower Cambrian Maotianshan shales of China 518 to 490 mya and is thought to be a vertebrate, although this is not conclusively proven. The species M. fengjiaoa is 28 mm long and 6 mm high. It is among the oldest possible craniates, found in the lower Cambrian Chengjiang (). It appears to have a skull and skeletal structures made of cartilage. There is no sign of biomineralization of the skeletal elements. The holotype was found in the Yuanshan member of the Qiongzhusi Formation in the Eoredlichia Zone near Haikou at Ercaicun, Kunming City, Yunnan, China. Only one species is known – ''Myllokunmingia fengjiaoa'' (Shu, Zhang & Han). Related creatures are ''Haikouichthys'' and '' Zhongjianichthys''. The animal has a distinct head and trunk with a forward sail-like (1.5 mm) dorsal fin and a ventral finfold (probably paired) further back. The maximum height of M. fengijaoa is at 6mm. The maximum height point i ...
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Myllokunmingia Fengjiaoa
''Myllokunmingia'' is a genus of basal chordate from the Lower Cambrian Maotianshan shales of China 518 to 490 mya and is thought to be a vertebrate, although this is not conclusively proven. The species M. fengjiaoa is 28 mm long and 6 mm high. It is among the oldest possible craniates, found in the lower Cambrian Chengjiang (). It appears to have a skull and skeletal structures made of cartilage. There is no sign of biomineralization of the skeletal elements. The holotype was found in the Yuanshan member of the Qiongzhusi Formation in the Eoredlichia Zone near Haikou at Ercaicun, Kunming City, Yunnan, China. Only one species is known – ''Myllokunmingia fengjiaoa'' (Shu, Zhang & Han). Related creatures are ''Haikouichthys'' and '' Zhongjianichthys''. The animal has a distinct head and trunk with a forward sail-like (1.5 mm) dorsal fin and a ventral finfold (probably paired) further back. The maximum height of M. fengijaoa is at 6mm. The maximum height poin ...
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Chordate
A chordate () is an animal of the phylum Chordata (). All chordates possess, at some point during their larval or adult stages, five synapomorphies, or primary physical characteristics, that distinguish them from all the other taxa. These five synapomorphies include a notochord, dorsal hollow nerve cord, endostyle or thyroid, pharyngeal slits, and a post-anal tail. The name “chordate” comes from the first of these synapomorphies, the notochord, which plays a significant role in chordate structure and movement. Chordates are also Bilateral symmetry, bilaterally symmetric, have a coelom, possess a circulatory system, and exhibit Metameric, metameric segmentation. In addition to the morphological characteristics used to define chordates, analysis of genome sequences has identified two conserved signature indels (CSIs) in their proteins: cyclophilin-like protein and mitochondrial inner membrane protease ATP23, which are exclusively shared by all vertebrates, tunicates and cep ...
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Vertebrate
Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, with currently about 69,963 species described. Vertebrates comprise such groups as the following: * jawless fish, which include hagfish and lampreys * jawed vertebrates, which include: ** cartilaginous fish (sharks, rays, and ratfish) ** bony vertebrates, which include: *** ray-fins (the majority of living bony fish) *** lobe-fins, which include: **** coelacanths and lungfish **** tetrapods (limbed vertebrates) Extant vertebrates range in size from the frog species ''Paedophryne amauensis'', at as little as , to the blue whale, at up to . Vertebrates make up less than five percent of all described animal species; the rest are invertebrates, which lack vertebral columns. The vertebrates traditionally include the hagfish, which do no ...
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Craniate
A craniate is a member of the Craniata (sometimes called the Craniota), a proposed clade of chordate animals with a skull of hard bone or cartilage. Living representatives are the Myxini (hagfishes), Hyperoartia (including lampreys), and the much more numerous Gnathostomata (jawed vertebrates).Campbell & Reece 2005 p. 676 Formerly distinct from vertebrates by including hagfish, molecular and anatomical research in the 21st century has led to the reinclusion of hagfish as vertebrates, making living craniates synonymous with living vertebrates. The clade was conceived largely on the basis of the Hyperoartia (lampreys and kin) being more closely related to the Gnathostomata (jawed vertebrates) than the Myxini (hagfishes). This, combined with an apparent lack of vertebral elements within the Myxini, suggested that the Myxini were descended from a more ancient lineage than the vertebrates, and that the skull developed before the vertebral column. The clade was thus composed of the Myx ...
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Haikouichthys
''Haikouichthys'' is an extinct genus of craniate (animals with notochords and distinct heads) that lived 518 million years ago, during the Cambrian explosion of multicellular life. ''Haikouichthys'' had a defined skull and other characteristics that have led paleontologists to label it a true craniate, and even to be popularly characterized as one of the Prehistoric fish, earliest fishes. Cladistics, Cladistic analysis indicates that the animal is probably a basal chordate or a basal craniate; but it does not possess sufficient features to be included uncontroversially even in either stem group. It was formally described in 1999. Description ''Haikouichthys'' is about long and is narrower than ''Myllokunmingia'', another Wiktionary:putative#English, putative chordate that comes from the same beds. The holotype of ''Haikouichthys ercaicunensis'' was found in the Yuansshan member of the Maotianshan Shales, Qiongzhusi Formation in the 'Eoredlichia' Zone near Haikou at Ercaicun, ...
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Maotianshan Shales
The Maotianshan Shales are a series of Cambrian, Early Cambrian deposits in the Chiungchussu Formation, famous for their ''Lagerstätte, Konservat Lagerstätten'', deposits known for the exceptional preservation of fossilized organisms or traces. The Maotianshan Shales form one of some forty Cambrian fossil locations worldwide exhibiting exquisite preservation of rarely preserved, non-mineralized soft tissue, comparable to the fossils of the Burgess Shale. They take their name from Maotianshan Hill (, Literal meaning: Hat Sky Mountain) in Chengjiang County, Yunnan Province, China. The most famous assemblage of organisms are referred to as the Chengjiang biota for the multiple scattered fossil sites in Chengjiang. The age of the Chengjiang Lagerstätte is locally termed Qiongzhusian, a stage correlated to the late Atdabanian Stage in Siberian sequences of the middle of the Early Cambrian. The shales date to ≤. The shales also contain the slightly younger Guanshan biota from Mal ...
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Haikouella
''Haikouella'' is an agnathan chordate from the Lower Cambrian Maotianshan Shales of Chengjiang County in Yunnan Province, China. An analysis in 2015 placed ''Haikouella'' as a junior synonym of ''Yunnanozoon,'' another Maotianshan shale Cambrian chordate''.'' It is similar to the form ''Yunnanozoon'', which is possibly a hemichordate. Still, there are anatomical differences from ''Yunnanozoon'', including a larger stomach and smaller (0.1 mm) pharyngeal teeth. ''Haikouella'' does not have bones or a movable jaw, but it otherwise resembles vertebrates. ''Haikouichthys'' and ''Myllokunmingia'', which seem to share significant fish-like characters, have been found in the same beds. Suspected hemichordates are also known from these deposits as well as from the Middle Cambrian Burgess Shale of British Columbia. Other than possible fish scales/plates from the Upper Cambrian of Wyoming, these Chinese fish-like chordates are one of the only known pre-Ordovician craniates. ''Haikou ...
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Notochord
In anatomy, the notochord is a flexible rod which is similar in structure to the stiffer cartilage. If a species has a notochord at any stage of its life cycle (along with 4 other features), it is, by definition, a chordate. The notochord consists of inner, vacuolated cells covered by fibrous and elastic sheaths, lies along the anteroposterior axis (''front to back''), is usually closer to the dorsal than the ventral surface of the embryo, and is composed of cells derived from the mesoderm. The most commonly cited functions of the notochord are: as a midline tissue that provides directional signals to surrounding tissue during development, as a skeletal (structural) element, and as a vertebral precursor. In lancelets the notochord persists throughout life as the main structural support of the body. In tunicates the notochord is present only in the larval stage, being completely absent in the adult animal. In these invertebrate chordates, the notochord is not vacuolated. In all ...
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Zhongjianichthys
''Zhongjianichthys rostratus'' is an extinct basal chordate that lived in the Cambrian period, approximately 518 million years ago. It is sometimes regarded as an early fish, and therefore as one of the first vertebrates. ''Zhongjianichthys'' is named after Zhongjian in China. Physical characteristics The eyes are located behind the antero-dorsal lobe and the mouth apparently did not have a jaw. It lacked scales, and had a thick skin. It is known to have had a thicker skin than other chordates at that time because, unlike ''Myllokunmingia,'' no impressions of its myomeres have been found. The ventral fin is low in height, and runs much of the body's length. Although ''Zhonjianichthys'' appears more advanced than its relatives due to the back-positioned eyes and the thicker skin, this may be convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous s ...
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Skull
The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, these two parts are the neurocranium and the viscerocranium ( facial skeleton) that includes the mandible as its largest bone. The skull forms the anterior-most portion of the skeleton and is a product of cephalisation—housing the brain, and several sensory structures such as the eyes, ears, nose, and mouth. In humans these sensory structures are part of the facial skeleton. Functions of the skull include protection of the brain, fixing the distance between the eyes to allow stereoscopic vision, and fixing the position of the ears to enable sound localisation of the direction and distance of sounds. In some animals, such as horned ungulates (mammals with hooves), the skull also has a defensive function by providing the mount (on the front ...
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Myomere
Myomeres are blocks of skeletal muscle tissue arranged in sequence, commonly found in aquatic chordates. Myomeres are separated from adjacent myomeres by connective fascia (myosepta) and most easily seen in larval fishes or in the olm. Myomere counts are sometimes used for identifying specimens, since their number corresponds to the number of vertebrae in the adults. Location varies, with some species containing these only near the tails, while some have them located near the scapular or pelvic girdles. Depending on the species, myomeres could be arranged in an epaxial or hypaxial manner. Hypaxial refers to ventral muscles and related structures while epaxial refers to more dorsal muscles. The horizontal septum divides these two regions in vertebrates from cyclostomes to gnathostomes. In terrestrial chordates, the myomeres become fused as well as indistinct, due to the disappearance of myosepta. Shape The shape of myomeres varies by species. Myomeres are commonly zig-zag, " ...
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Cambrian Fish
The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Cambrian biolo ...
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