Byronosaurus
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Byronosaurus
''Byronosaurus'' is a genus of troodontid dinosaur from the Late Cretaceous Period of Mongolia. Discovery and naming In 1993, Michael Novacek, a member of an American Museum of Natural History expedition to the Gobi Desert, discovered the skeleton of a small theropod at Ukhaa Tolgod. This was further excavated in 1994 and 1995. The find was illustrated in a publication in 1994. On 15 July 1996, at the Bolor's Hill site, about eight kilometers (five miles) away from the original location, a second specimen was discovered, a skull. In 2000, Mark Norell, Peter Makovicky and James Clark named and described the type species ''Byronosaurus jaffei''. The species name as a whole honoured Byron Jaffe, "in recognition of his family's support for the Mongolian Academy of Sciences-American Museum of Natural History Paleontological Expeditions". The holotype, IGM 100/983, was found in a layer of the Djadochta Formation dating from the late Campanian. It consists of a partial skeleton wi ...
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Byronosaurus
''Byronosaurus'' is a genus of troodontid dinosaur from the Late Cretaceous Period of Mongolia. Discovery and naming In 1993, Michael Novacek, a member of an American Museum of Natural History expedition to the Gobi Desert, discovered the skeleton of a small theropod at Ukhaa Tolgod. This was further excavated in 1994 and 1995. The find was illustrated in a publication in 1994. On 15 July 1996, at the Bolor's Hill site, about eight kilometers (five miles) away from the original location, a second specimen was discovered, a skull. In 2000, Mark Norell, Peter Makovicky and James Clark named and described the type species ''Byronosaurus jaffei''. The species name as a whole honoured Byron Jaffe, "in recognition of his family's support for the Mongolian Academy of Sciences-American Museum of Natural History Paleontological Expeditions". The holotype, IGM 100/983, was found in a layer of the Djadochta Formation dating from the late Campanian. It consists of a partial skeleton wi ...
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Xixiasaurus Henanensis
''Xixiasaurus'' () is a genus of troodontid dinosaur that lived during the Late Cretaceous Period in what is now China. The only known specimen was discovered in Xixia County, Henan Province, in central China, and became the holotype of the new genus and species ''Xixiasaurus henanensis'' in 2010. The names refer to the areas of discovery, and can be translated as "Henan Xixia lizard". The specimen consists of an almost complete skull (except for the hindmost portion), part of the lower jaw, and teeth, as well as a partial right forelimb. ''Xixiasaurus'' is estimated to have been long and to have weighed . As a troodontid, it would have been bird-like and lightly built, with grasping hands and an enlarged sickle-shaped claw on the second toe. Its skull was long, with a long, low snout that formed a tapering U-shape when seen from below. The of the forehead was dome-like in side view, which indicates it had an enlarged . It differed from other troodontids in that the front of ...
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Xixiasaurus
''Xixiasaurus'' () is a genus of troodontid dinosaur that lived during the Late Cretaceous Period in what is now China. The only known specimen was discovered in Xixia County, Henan Province, in central China, and became the holotype of the new genus and species ''Xixiasaurus henanensis'' in 2010. The names refer to the areas of discovery, and can be translated as "Henan Xixia lizard". The specimen consists of an almost complete skull (except for the hindmost portion), part of the lower jaw, and teeth, as well as a partial right forelimb. ''Xixiasaurus'' is estimated to have been long and to have weighed . As a troodontid, it would have been bird-like and lightly built, with grasping hands and an enlarged sickle-shaped claw on the second toe. Its skull was long, with a long, low snout that formed a tapering U-shape when seen from below. The of the forehead was dome-like in side view, which indicates it had an enlarged . It differed from other troodontids in that the front ...
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Troodontidae
Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil discoveries of complete and articulated specimens (including specimens which preserve feathers, eggs, embryos, and complete juveniles), have helped to increase understanding about this group. Anatomical studies, particularly studies of the most primitive troodontids, like ''Sinovenator'', demonstrate striking anatomical similarities with ''Archaeopteryx'' and primitive dromaeosaurids, and demonstrate that they are relatives comprising a clade called Paraves. Description Troodontids are a group of small, bird-like, gracile maniraptorans. All troodontids have unique features of the skull, such as large numbers of closely spaced teeth in the lower jaw. Troodontids have sickle-claws and raptorial hands, and some of the highest non-avian encephali ...
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Djadochta Formation
The Djadochta Formation (sometimes Transcription (linguistics), transcribed and also known as Djadokhta, Djadokata, or Dzhadokhtskaya) is a highly fossiliferous geological formation situated in Central Asia, Gobi Desert, dating from the Late Cretaceous Period (geology), period, about 75 million to 71 million years ago. The type locality (geology), type locality is the Bayn Dzak locality, famously known as the Flaming Cliffs. Dinosaur, mammal, and other reptile remains are among the fossils that have been recovered from the formation. Excavation history The Djadochta Formation was first documented and explored—though only a single locality—during paleontological expeditions of the American Museum of Natural History in 1922–1925, which were part of the Central Asiatic Expeditions. The expeditions were led by Roy Chapman Andrews, in company of Walter Willis Granger as chief paleontologist and field team. The team did extensive exploration at the Bayn Dzak (formerly Shabarakh Us ...
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Mark Norell
Mark Allen Norell (born July 26, 1957) is an American paleontologist, acknowledged as one of the most important living vertebrate paleontologists. He is currently the chairman of paleontology and a research associate at the American Museum of Natural History. He is best known as the discoverer of the first theropod embryo and for the description of feathered dinosaurs. Norell is credited with the naming of the genera ''Apsaravis'', ''Byronosaurus'', '' Citipati'', '' Tsaagan'', and '' Achillobator''. His work regularly appears in major scientific journals (including cover stories in Science and Nature) and was listed by Time magazine as one of the ten most significant science stories of 1993, 1994 and 1996. Norell is both a fellow of the Explorer's Club and the Willi Hennig Society. Career Norell's research has encompassed a number of different areas, from the theoretical study of diversity through time, his doctoral dissertation on alligator phylogeny, and his postdoctoral w ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia an ...
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Paratype
In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). Often there is more than one paratype. Paratypes are usually held in museum research collections. The exact meaning of the term ''paratype'' when it is used in zoology is not the same as the meaning when it is used in botany. In both cases however, this term is used in conjunction with ''holotype''. Zoology In zoological nomenclature, a paratype is officially defined as "Each specimen of a type series other than the holotype.", ''International Code of Zoological Nomenclature'' In turn, this definition relies on the definition of a "type series". A type series is the material (specimens of organisms) that was cited in the original publication of the new species or subspecies, and was not excluded from being type material by the author ( ...
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Maxilla
The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxillary bones are fused at the intermaxillary suture, forming the anterior nasal spine. This is similar to the mandible (lower jaw), which is also a fusion of two mandibular bones at the mandibular symphysis. The mandible is the movable part of the jaw. Structure In humans, the maxilla consists of: * The body of the maxilla * Four processes ** the zygomatic process ** the frontal process of maxilla ** the alveolar process ** the palatine process * three surfaces – anterior, posterior, medial * the Infraorbital foramen * the maxillary sinus * the incisive foramen Articulations Each maxilla articulates with nine bones: * two of the cranium: the frontal and ethmoid * seven of the face: the nasal, zygomatic, lacrimal ...
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Velociraptor
''Velociraptor'' (; ) is a genus of small dromaeosaurid dinosaur that lived in Asia during the Late Cretaceous epoch, about 75 million to 71 million years ago. Two species are currently recognized, although others have been assigned in the past. The type species is ''V. mongoliensis''; fossils of this species have been discovered in the Djadochta Formation, Mongolia. A second species, ''V. osmolskae'', was named in 2008 for skull material from the Bayan Mandahu Formation, China. Smaller than other dromaeosaurids like ''Deinonychus'' and ''Achillobator'', ''Velociraptor'' was about long with a body mass between . It nevertheless shared many of the same anatomical features. It was a bipedal, feathered carnivore with a long tail and an enlarged sickle-shaped claw on each hindfoot, which is thought to have been used to tackle and restrain prey. ''Velociraptor'' can be distinguished from other dromaeosaurids by its long and low skull, with an upturned snout. ''Velociraptor'' ...
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Plesiomorph
In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, and synapomorphy, all mean a trait shared between species because they share an ancestral species. Apomorphic and synapomorphic characteristics convey much information about evolutionary clades and can be used to define taxa. However, plesiomorphic and symplesiomorphic characteristics cannot. The term ''symplesiomorphy'' was introduced in 1950 by German entomologist Willi Hennig. Examples A backbone is a plesiomorphic trait shared by birds and mammals, and does not help in placing an animal in one or the other of these two clades. Birds and mammals share this trait because both clades are descended from the same far distant ancestor. Other clades, e.g. snakes, lizards, turtles, fish, frogs, all have backbones and none are either birds ...
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Almas Ukhaa
''Almas'' is a genus of troodontid theropod dinosaur from the Late Cretaceous of Mongolia. It contains a single species, ''Almas ukhaa'', named in 2017 by Pei Rui and colleagues, based on a partial articulated skeleton. The only known specimen was found in the Djadochta Formation, which is late Campanian in age. Discovery and naming In 1993, a joint expedition by the American Museum of Natural History and the Mongolian Academy of Sciences discovered near Ukhaa Tolgod, the Flaming Cliffs, a skeleton of a small theropod. It was prepared by Amy Davidson. Though in subsequent years its traits were inserted in some data matrices of phylogenetic analyses, a description of the fossil was never published. In 2017, the type species ''Almas ukhaa'' was named and described by Pei Rui, Mark Norell, Daniel Barta, Gabriel Bever, Michael Pittman and Xu Xing. The generic name refers to the almas, "wild man" in Mongolian, a man-like creature from Mongolian folklore. The specific name refers ...
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